"The science of rudimentary organs," continues Mr. Wallace, "which Haeckel terms 'dysteleology, or the doctrine of purposelessness,' is here discussed, and a number of interesting examples are given, the conclusion being that they prove the mechanical or monistic conception of the origin of organisms to be correct, and the idea of any 'all-wise creative plan an ancient fable.'" I see no reason to suppose, or again not to suppose, an all-wise creative plan. I decline to go into this question, believing it to be not yet ripe, nor nearly ripe, for consideration. I see purpose, however, in rudimentary organs as much as in useful ones, but a spent or extinct purpose—a purpose which has been fulfilled, and is now forgotten—the rudimentary organ being repeated from force of habit, indolence, and dislike of change, so long as it does not, to use the words of Buffon, "stand in the way of the fair development" of other parts which are found useful and necessary. I demur, therefore, to the inference of "purposelessness" which I gather that Professor Haeckel draws from these organs.
In the 'Academy' for April 19, 1879, Mr. Wallace quotes Professor Haeckel as saying that our "highly purposive and admirably-constituted sense-organs have developed without premeditated aim; that they have originated by the same mechanical process of Natural Selection, by the same constant interaction of Adaptation and Heredity [what is Heredity but another word for unknown causes, unless it is explained in some such manner as in 'Life and Habit'?] by which all the other purposive contrivances of the animal organization have been slowly and gradually evolved during the struggle for existence."
I see no evidence for "premeditated aim" at any modification very far in advance of an existing organ, any more than I do for "premeditated aim" on man's part at any as yet inconceivable mechanical invention; but as in the case of man's inventions, so also in that of the organs of animals and plants, modification is due to the accumulation of small, well-considered improvements, as found necessary in practice, and the conduct of their affairs. Each step having been purposive, the whole road has been travelled purposively; nor is the purposiveness of such an organ, we will say, as the eye, barred by the fact that invention has doubtless been aided by some of those happy accidents which from time to time happen to all who keep their wits about them, and know how to turn the gifts of Fortune to account.
FOOTNOTES:
[371] 'Origin of Species,' p. 109.
[372] 'Origin of Species, p. 401.
[373] 'Phil. Zool.,' tom. i. p. 242.
[374] 'Phil. Zool.,' tom. i. p. 244.
[375] 'Phil. Zool.,' tom. i. p. 245.