If the primordial cell had been only capable of reproducing itself once, there would have followed a single line of descendants, the chain of which might at any moment have been broken by casualty. Doubtless the millionth repetition would have differed very materially from the original—as widely, perhaps, as we differ from the primordial cell; but it would only have differed by addition, and could no more in any generation resume its latest development without having passed through the initial stage of being what its first forefather was, and doing what its first forefather did, and without going through all or a sufficient number of the steps whereby it had reached its latest differentiation, than water can rise above its own level.

The very idea, then, of reproduction involves, unless I am mistaken, that, no matter how much the creature reproducing itself may gain in power and versatility, it must still always begin with itself again in each generation. The primordial cell being capable of reproducing itself not only once, but many times over, each of the creatures which it produces must be similarly gifted; hence the geometrical ratio of increase and the existing divergence of type. In each generation it will pass rapidly and unconsciously through all the earlier stages of which there has been infinite experience, and for which the conditions are reproduced with sufficient similarity to cause no failure of memory or hesitation; but in each generation, when it comes to the part in which the course is not so clear, it will become conscious; still, however, where the course is plain, as in breathing, digesting, &c., retaining unconsciousness. Thus organs which present all the appearance of being designed—as, for example, the tip for its beak prepared by the embryo chicken—would be prepared in the end, as it were, by rote, and without sense of design, though none the less owing their origin to design.

The question is not concerning evolution, but as to the main cause which has led to evolution in such and such shapes. To me it seems that the “Origin of Variation,” whatever it is, is the only true “Origin of Species,” and that this must, as Lamarck insisted, be looked for in the needs and experiences of the creatures varying. Unless we can explain the origin of variations, we are met by the unexplained at every step in the progress of a creature from its original homogeneous condition to its differentiation, we will say, as an elephant; so that to say that an elephant has become an elephant through the accumulation of a vast number of small, fortuitous, but unexplained, variations in some lower creatures, is really to say that it has become an elephant owing to a series of causes about which we know nothing whatever, or, in other words, that one does not know how it came to be an elephant. But to say that an elephant has become an elephant owing to a series of variations, nine-tenths of which were caused by the wishes of the creature or creatures from which the elephant is descended—this is to offer a reason, and definitely put the insoluble one step further back. The question will then turn upon the sufficiency of the reason—that is to say, whether the hypothesis is borne out by facts.

The effects of competition would, of course, have an extremely important effect upon any creature, in the same way as any other condition of nature under which it lived, must affect its sense of need and its opinions generally. The results of competition would be, as it were, the decisions of an arbiter settling the question whether such and such variation was really to the animal’s advantage or not—a matter on which the animal will, on the whole, have formed a pretty fair judgement for itself. Undoubtedly the past decisions of such an arbiter would affect the conduct of the creature, which would have doubtless had its shortcomings and blunders, and would amend them. The creature would shape its course according to its experience of the common course of events, but it would be continually trying and often successfully, to evade the law by all manner of sharp practice. New precedents would thus arise, so that the law would shift with time and circumstances; but the law would not otherwise direct the channels into which life would flow, than as laws, whether natural or artificial, have affected the development of the widely differing trades and professions among mankind. These have had their origin rather in the needs and experiences of mankind than in any laws.

To put much the same as the above in different words. Assume that small favourable variations are preserved more commonly, in proportion to their numbers, than is perhaps the case, and assume that considerable variations occur more rarely than they probably do occur, how account for any variation at all? “Natural selection” cannot create the smallest variation unless it acts through perception of its mode of operation, recognised inarticulately, but none the less clearly, by the creature varying. “Natural selection” operates on what it finds, and not on what it has made. Animals that have been wise and lucky live longer and breed more than others less wise and lucky. Assuredly. The wise and lucky animals transmit their wisdom and luck. Assuredly. They add to their powers, and diverge into widely different directions. Assuredly. What is the cause of this? Surely the fact that they were capable of feeling needs, and that they differed in their needs and manner of gratifying them, and that they continued to live in successive generations, rather than the fact that when lucky and wise they thrived and bred more descendants. This last is an accessory hardly less important for the development of species than the fact of the continuation of life at all; but it is an accessory of much the same kind as this, for if animals continue to live at all, they must live in some way, and will find that there are good ways and bad ways of living. An animal which discovers the good way will gradually develop further powers, and so species will get further and further apart; but the origin of this is to be looked for, not in the power which decides whether this or that way was good, but in the cause which determines the creature, consciously or unconsciously, to try this or that way.

But Mr. Darwin might say that this is not a fair way of stating the issue. He might say, “You beg the question; you assume that there is an inherent tendency in animals towards progressive development, whereas I say that there is no good evidence of any such tendency. I maintain that the differences that have from time to time arisen have come about mainly from causes so far beyond our ken, that we can only call them spontaneous; and if so, natural selection which you must allow to have at any rate played an important part in the accumulation of variations, must also be allowed to be the nearest thing to the cause of Specific differences, which we are able to arrive at.”

Thus he writes (“Natural Selection,” p. 176, ed. 1876): “Although we have no good evidence of the existence in organic beings of a tendency towards progressive development, yet this necessarily follows, as I have attempted to show in the fourth chapter, through the continued action of natural selection.” Mr. Darwin does not say that organic beings have no tendency to vary at all, but only that there is no good evidence that they have a tendency to progressive development, which, I take it, means, to see an ideal a long way off, and very different to their present selves, which ideal they think will suit them, and towards which they accordingly make. I would admit this as contrary to all experience. I doubt whether plants and animals have any innate tendency to vary at all, being led to question this by gathering from “Plants and Animals under Domestication” that this is Mr. Darwin’s own opinion. I am inclined rather to think that they have only an innate power to vary slightly, in accordance with changed conditions, and an innate capability of being affected both in structure and instinct, by causes similar to those which we observe to affect ourselves. But however this may be, they do vary somewhat, and unless they did, they would not in time have come to be so widely different from each other as they now are. The question is as to the origin and character of these variations.

We say they mainly originate in a creature through a sense of its needs, and vary through the varying surroundings which will cause those needs to vary, and through the opening up of new desires in many creatures, as the consequence of the gratification of old ones; they depend greatly on differences of individual capacity and temperament; they are communicated, and in the course of time transmitted, as what we call hereditary habits or structures, though these are only, in truth, intense and epitomised memories of how certain creatures liked to deal with protoplasm. The question whether this or that is really good or ill, is settled, as the proof of the pudding by the eating thereof, i.e., by the rigorous competitive examinations through which most living organisms must pass. Mr. Darwin says that there is no good evidence in support of any great principle, or tendency on the part of the creature itself, which would steer variation, as it were, and keep its head straight, but that the most marvellous adaptations of structures to needs are simply the result of small and blind variations, accumulated by the operation of “natural selection,” which is thus the main cause of the origin of species.

Enough has perhaps already been said to make the reader feel that the question wants reopening; I shall, therefore, here only remark that we may assume no fundamental difference as regards intelligence, memory, and sense of needs to exist between man and the lowest animals, and that in man we do distinctly see a tendency towards progressive development, operating through his power of profiting by and transmitting his experience, but operating in directions which man cannot foresee for any long distance. We also see this in many of the higher animals under domestication, as with horses which have learnt to canter and dogs which point; more especially we observe it along the line of latest development, where equilibrium of settled convictions has not yet been fully attained. One neither finds nor expects much a priori knowledge, whether in man or beast; but one does find some little in the beginnings of, and throughout the development of, every habit, at the commencement of which, and on every successive improvement in which, deductive and inductive methods are, as it were, fused. Thus the effect, where we can best watch its causes, seems mainly produced by a desire for a definite object—in some cases a serious and sensible desire, in others an idle one, in others, again, a mistaken one; and sometimes by a blunder which, in the hands of an otherwise able creature, has turned up trumps. In wild animals and plants the divergences have been accumulated, if they answered to the prolonged desires of the creature itself, and if these desires were to its true ultimate good; with plants or animals under domestication they have been accumulated if they answered a little to the original wishes of the creature, and much, to the wishes of man. As long as man continued to like them, they would be advantageous to the creature; when he tired of them, they would be disadvantageous to it, and would accumulate no longer. Surely the results produced in the adaptation of structure to need among many plants and insects are better accounted for on this, which I suppose to be Lamarck’s view, namely, by supposing that what goes on amongst ourselves has gone on amongst all creatures, than by supposing that these adaptations are the results of perfectly blind and unintelligent variations.

Let me give two examples of such adaptations, taken from Mr. St. George Mivart’s “Genesis of Species,” to which work I would wish particularly to call the reader’s attention. He should also read Mr. Darwin’s answers to Mr. Mivart (p. 176, “Natural Selection,” ed. 1876, and onwards).