The proportion of the food of oxen, sheep, and pigs, which is consumed in maintaining their vital functions, has not been accurately ascertained; probably, as in the case of man, it is strictly proportionate to the animal's weight. We can determine the amount of plastic food consumed by an animal during a given period: we can ascertain the increase (if any) in the weight of its body; and finally, we can weigh and analyse its egesta. With these data it is comparatively easy to ascertain the quantity of food which produced the increase in the animal's weight; but they do not enable us to determine the amount expended in keeping it alive, because the egesta might be largely made up of unappropriated food—organised matter which had done no work in the animal body. When we come to know the precise quantity of nitrogen, in a purely, or nearly pure, mineral form[!--14--][14] excreted by an animal, then we shall be in a position to estimate the proportion of its food expended in sustaining the essential vital processes which continuously go on in its body. But although we are in ignorance as to the precise quantity of flesh-formers expended in keeping the animal alive, we know pretty accurately the amount which is consumed in producing a given weight of its flesh, or rather in causing a certain increase in its weight. This knowledge is the result of numerous investigations, of which by far the most valuable are those of Lawes and Gilbert. These experimenters found that fattening pigs stored up about 7½ per cent. of the plastic materials of their food, whilst sheep accumulated somewhat less than 5 per cent. That is, 92½ out of every 100 lbs. weight of the nitrogenous food of the pig, and 95 out of every 100 lbs. of that of the sheep, are eliminated in the excretions of those animals.

It appears from the results of Lawes and Gilbert's experiments, that pigs store up in their increase about 20 per cent., sheep 12 per cent., and oxen 8 per cent. of their (dry) food. The relative increase of the fatty, nitrogenous, and mineral constituents whilst fattening, are shown in this table.

Cases.Estimated per cent. in Increase whilst Fattening.
Mineral matter (ash.)Nitrogenous matter (dry). Fat (dry). Total dry substance.
Average of 98 oxen 1·47 7·69 66·2 75·4
Average of 348 sheep 1·80 7·13 70·4 79·53
Average of 80 pigs 0·44 6·44 71·5 78·40

The quantity of food consumed daily by an animal is, as might be expected, proportionate to the weight of its body. The pig consumes, for every 100 lbs. of its weight, from 26 to 30 lbs. of food, the sheep 15 lbs., and the ox 12 to 13 lbs. These figures and the statements which I have made relative to the proportions of fat and plastic elements in the animals' bodies, apply to them in their fattening state, and when the food is of a highly nutritious character. The calf and the young pig will make use—to cause their increase—of a larger portion of nitrogenous matters. The sheep, however, being early brought to maturity, will, even when very young, store up the plastic and non-plastic constituents of its food, in nearly the same relative proportions that I have mentioned.

As it is the food taken into the body that produces heat and motion, it might at first sight appear an easy matter to determine the amount of heat or of motion which a given weight of a particular kind of food is capable of producing within the animal mechanism. But this performance is not so easy a task as it appears to be. In the first place, all of the food may not be perfectly oxidised, though thoroughly disorganised within the body; secondly, as animals rarely subsist on one kind of food, it is difficult, when they are supplied with mixed aliments, to determine which of them is the most perfectly decomposed. But though the difficulties which I have mentioned, and many others, render the task of determining the nutritive values of food substances difficult, the problem is by no means insoluble, and, in fact, is in a fair way of being solved. Professor Frankland, in a paper published in the number of the Philosophical Magazine for September, 1866, determines the relative alimental value of foods by ascertaining the quantity of heat evolved by each when burned in oxygen gas. From the results of these researches he has constructed a table, showing the amount of food necessary to keep a man alive for twenty-four hours. The following figures, which I select from this table, are of interest to the stock-feeder:—

Weight necessary to sustain a man's life
for twenty-four hours.
Kinds of Food. Ounces.
Potatoes 13·4
Apples 20·7
Oatmeal 3·4
Flour 3·5
Pea Meal 3·5
Bread 6·4
Milk 21·2
Carrots 25·6
Cabbage 31·8
Butter 1·8
Lump Sugar 3·9

These figures show the relative calefacient, or heat-producing powers of the different foods named outside the body; but there is some doubt as to their having the same relative values when burned within the body. The woody fibre of the carrots and cabbages is very combustible in the coal furnace, but it is very doubtful if more than 20 or 30 per cent. of this substance is ever burned in the animal furnace. However, such inquiries as those carried out by Frankland possess great value; and tables constructed upon their results cannot fail to be useful in the drawing up of dietary scales, whether for man or for the inferior animals.

I may here remark, that in my opinion the nutritive value of food admits of being very accurately determined by the adoption of the following method:—