By the hypothesis of such a slow and gradual subsidence of the land we may account for the fact that almost everywhere in N. America and Northern Europe the boulder formation rests on a polished and furrowed surface of rock,—a fact by no means obliging us to imagine, as some think, that the polishing and grooving action was, as a whole, anterior in date to the transportation of the erratics. During the successive depression of high land, varying originally in height from 1000 to 3000 feet above the sea-level, every portion of the surface would be brought down by turns to the level of the ocean, so as to be converted first into a coast-line, and then into a shoal; and at length, after being well scored by the stranding upon it of thousands of icebergs, might be sunk to a depth of several hundred fathoms. By the constant depression of land, the coast would recede farther and farther from the successively formed zones of polished and striated rock, each outer zone becoming in its turn so deep under water as to be no longer grated upon by the heaviest icebergs. Such sunken areas would then simply serve as receptacles of mud, sand, and boulders dropped from melting ice, perhaps to a depth scarcely, if at all, inhabited by testacea and zoophytes. Meanwhile, during the formation of the unstratified and unfossiliferous mass in deeper water, the smoothing and furrowing of shoals and beaches is still going on elsewhere upon and near the coast in full activity. If at length the subsidence should cease, and the direction of the movement of the earth's crust be reversed, the sunken area covered with drift would be slowly reconverted into land. The boulder deposit, before emerging, would then for a time be brought within the action of the waves, tides, and currents, so that its upper portion, being partially disturbed, would have its materials re-arranged and stratified. Streams also flowing from the land would in some places throw down layers of sediment upon the till. In that case, the order of superposition will be, first and uppermost, sand, loam, and gravel occasionally fossiliferous; secondly, an unstratified and unfossiliferous mass, for the most part of much older date than the preceding, with angular erratics, or with boulders interspersed; and, thirdly, beneath the whole, a surface of polished and furrowed rock. Such a succession of events seems to have prevailed very widely on both sides of the Atlantic, the travelled blocks having been carried in general from the North Pole southwards, but mountain chains having in some cases served as independent centres of dispersion, of which the Alps present the most conspicuous example.

It is by no means rare to meet with boulders imbedded in drift which are worn flat on one or more of their sides, the surface being at the same time polished, furrowed, and striated. They may have been so shaped in a glacier before they reached the sea, or when they were fixed in the bottom of an iceberg as it ran aground. We learn from Mr. Charles Martins that the glaciers of Spitzbergen project from the coast into a sea between 100 and 400 feet deep; and that numbers of striated pebbles or blocks are there seen to disengage themselves from the overhanging masses of ice as they melt, so as to fall at once into deep water.[136-A]

That they should retain such markings when again upraised above the sea ought not to surprise us, when we remember that rippled sands, and the cracks in clay dried between high and low water, and the foot-tracks of animals and rain-drops impressed on mud, and other superficial markings, are all found fossil in rocks of various ages.

On the other hand, it is not difficult to account for the absence in many districts of striated and scored pebbles and boulders in glacial deposits, for they may have been exposed to the action of the waves on a coast while it was sinking beneath or rising above the sea. No shingle on an ordinary sea-beach exhibits such striæ, and at a very short distance from the termination of a glacier every stone in the bed of the torrent which gushes out from the melting ice is found to have lost its glacial markings by being rolled for a distance even of a few hundred yards.

The usual dearth of fossil shells in glacial clays well fitted to preserve organic remains may, perhaps, be owing, as already hinted, to the absence of testacea in the deep sea, where the undisturbed accumulation of boulders melted out of very large bergs may take place. In the Ægean and other parts of the Mediterranean, the zero of animal life, according to Prof. E. Forbes, is approached at a depth of about 300 fathoms. In tropical seas it would descend farther down, just as vegetation ascends higher on the mountains of hot countries. Near the pole, on the other hand, the same zero would be reached much sooner both on the hills and in the sea. If the ocean was filled with floating bergs, and a low temperature prevailed in the northern hemisphere during the glacial period, even the shallow part of the sea might have been uninhabitable, or very thinly peopled with living beings. It may also be remarked that the melting of ice in some fiords in Norway freshens the water so as to destroy marine life, and famines have been caused in Iceland by the stranding of icebergs drifted from the Greenland coast, which have required several years to melt, and have not only injured the hay harvest by cooling the atmosphere, but have driven away the fish from the shore by chilling and freshening the sea.

If the cold of the glacial epoch came on slowly, if it was long before it reached its greatest intensity, and again if it abated gradually, we may expect to find the earliest and latest formed drift less barren of organic remains than that deposited during the coldest period. We may also expect that along the southern limits of the drift during the whole glacial epoch, there would be an intimate association of transported matter of northern origin with fossil-bearing sediment, whether marine or freshwater, belonging to more southern seas, rivers, and continents.

That in the United States, the Mastodon giganteus was very abundant after the drift period is evident from the fact that entire skeletons of this animal are met with in bogs and lacustrine deposits occupying hollows in the drift. They sometimes occur in the bottom even of small ponds recently drained by the agriculturist for the sake of the shell marl. I examined one of these spots at Geneseo in the state of New York, from which the bones, skull, and tusk of a Mastodon had been procured in the marl below a layer of black peaty earth, and ascertained that all the associated freshwater and land shells were of a species now common in the same district. They consisted of several species of Lymnea, of Planorbis bicarinatus, Physa heterostropha, &c.

In 1845 no less than six skeletons of the same species of Mastodon were found in Warren County, New Jersey, 6 feet below the surface, by a farmer who was digging out the rich mud from a small pond which he had drained. Five of these skeletons were lying together, and a large part of the bones crumbled to pieces as soon as they were exposed to the air. But nearly the whole of the other skeleton, which lay about 10 feet apart from the rest, was preserved entire, and proved the correctness of Cuvier's conjecture respecting this extinct animal, namely, that it had twenty ribs like the living elephant. From the clay in the interior within the ribs, just where the contents of the stomach might naturally have been looked for, seven bushels of vegetable matter were extracted. I submitted some of this matter to Mr. A. Henfrey of London for microscopic examination, and he informs me that it consists of pieces of small twigs of a coniferous tree of the Cypress family, probably the young shoots of the white cedar, Thuja occidentalis, still a native of North America, on which therefore we may conclude that this extinct Mastodon once fed.

Another specimen of the same quadruped, the most complete and probably the largest ever found, was exhumed in 1845 in the town of Newburg, New York, the length of the skeleton being 25 feet, and its height 12 feet. The anchylosing of the last two ribs on the right side afforded Dr. John C. Warren a true gauge for the space occupied by the intervertebrate substance, so as to enable him to form a correct estimate of the entire length. The tusks when discovered were 10 feet long, but a part only could be preserved. The large proportion of animal matter in the tusk, teeth, and bones of some of these fossil mammalia is truly astonishing. It amounts in some cases, as Dr. C. T. Jackson has ascertained by analysis, to 27 per cent., so that when all the earthy ingredients are removed by acids, the form of the bone remains as perfect, and the mass of animal matter is almost as firm, as in a recent bone subjected to similar treatment.

It would be rash, however, to infer from such data that these quadrupeds were mired in modern times, unless we use that term strictly in a geological sense. I have shown that there is a fluviatile deposit in the valley of the Niagara, containing shells of the genera Melania, Lymnea, Planorbis, Valvata, Cyclas, Unio, and Helix, &c., all of recent species, from which the bones of the great Mastodon have been taken in a very perfect state. Yet the whole excavation of the ravine, for many miles below the Falls, has been slowly effected since that fluviatile deposit was thrown down.