England.—In a cave at Kirkdale, about twenty-five miles N.N.E. of York, the remains of about 300 hyænas, belonging to individuals of every age, have been detected. The species (Hyæna spelæa) is extinct, and was larger than the fierce Hyæna crocuta of South Africa, which it most resembled. Dr. Buckland, after carefully examining the spot, proved that the Hyænas must have lived there; a fact attested by the quantity of their dung, which, as in the case of the living hyæna, is of nearly the same composition as bone, and almost as durable. In the cave were found the remains of the ox, young elephant, hippopotamus, rhinoceros, horse, bear, wolf, hare, water-rat, and several birds. All the bones have the appearance of having been broken and gnawed by the teeth of the hyænas; and they occur confusedly mixed in loam or mud, or dispersed through a crust of stalagmite which covers it. In these and many other cases it is supposed that portions of herbivorous quadrupeds have been dragged into caverns by beasts of prey, and have served as their food, an opinion quite consistent with the known habits of the living hyæna.
No less than thirty-seven species of mammalia are enumerated by Professor Owen as having been discovered in the caves of the British islands, of which eighteen appear to be extinct, while the others still survive in Europe. They were not washed to the spots where the fossils now occur by a great flood; but lived and died, one generation after another, in the places where they lie buried. Among other arguments in favour of this conclusion may be mentioned the great numbers of the shed antlers of deer discovered in caves and in freshwater strata throughout England.[155-A]
Examples also occur of fissures into which animals have fallen from time to time, or have been washed in from above, together with alluvial matter and fragments of rock detached by frost, forming a mass which may be united into a bony breccia by stalagmitic infiltrations. Frequently we discover a long suite of caverns connected by narrow and irregular galleries, which hold a tortuous course through the interior of mountains, and seem to have served as the subterranean channels of springs and engulphed rivers. Many streams in the Morea are now carrying bones, pebbles, and mud into underground passages of this kind.[155-B] If, at some future period, the form of that country should be wholly altered by subterranean movements and new valleys shaped out by denudation, many portions of the former channels of these engulphed streams may communicate with the surface, and become the dens of wild beasts, or the recesses to which quadrupeds retreat to die. Certain caves of France, Germany, and Belgium, may have passed successively through these different conditions, and in their last state may have remained open to the day for several tertiary periods. It is nevertheless remarkable, that on the continent of Europe, as in England, the fossil remains of mammalia belong almost exclusively to those of the Newer Pliocene and Post-Pliocene periods, and not to the Miocene or Eocene epochs, and when they are accompanied by land or river shells, these agree in great part, or entirely, with recent species.
As the preservation of the fossil bones is due to a slow and constant supply of stalactite, brought into the caverns by water dropping from the roof, the source and origin of this deposit has been a subject of curious inquiry. The following explanation of the phenomenon has been recently suggested by the eminent chemist Liebig. On the surface of Franconia, where the limestone abounds in caverns, is a fertile soil, in which vegetable matter is continually decaying. This mould or humus, being acted on by moisture and air, evolves carbonic acid which is dissolved by rain. The rain water, thus impregnated, permeates the porous limestone, dissolves a portion of it, and afterwards, when the excess of carbonic acid evaporates in the caverns, parts with the calcareous matter, and forms stalactite.
Australian cave-breccias.—Ossiferous breccias are not confined to Europe, but occur in all parts of the globe; and those lately discovered in fissures and caverns in Australia correspond closely in character with what has been called the bony breccia of the Mediterranean, in which the fragments of bone and rock are firmly bound together by a red ochreous cement.
Some of these caves have been examined by Sir T. Mitchell in the Wellington Valley, about 210 miles west of Sidney, on the river Bell, one of the principal sources of the Macquarie, and on the Macquarie itself. The caverns often branch off in different directions through the rock, widening and contracting their dimensions, and the roofs and floors are covered with stalactite. The bones are often broken, but do not seem to be water-worn. In some places they lie imbedded in loose earth, but they are usually included in a breccia.
The remains found most abundantly are those of the kangaroo, of which there are four species, besides which the genera Hypsiprymnus, Phalangista, Phascolomys, and Dasyurus, occur. There are also bones, formerly conjectured by some osteologists to belong to the hippopotamus, and by others to the dugong, but which are now referred by Mr. Owen to a marsupial genus, allied to the Wombat.
Fig. 126.
