The total number of mollusca from the faluns, in my possession, is 302, of which 45 only were found by Mr. Wood to be common to the Suffolk crag. The number of corals obtained by me at Doué, and other localities before adverted to, amounts to 43, as determined by Mr. Lonsdale, of which 7 agree specifically with those of the Suffolk crag. Only one has, as yet, been identified with a living species. But it is difficult, if not impossible, to institute at present a satisfactory comparison between fossil and recent Polyparia, from the deficiency of our knowledge of the living species. Some of the genera occurring fossil in Touraine, as the Astrea, Lunulites, and Dendrophyllia, have not been found in European seas north of the Mediterranean; nevertheless the Polyparia of the faluns do not seem to indicate on the whole so warm a climate as would be inferred from the shells.
It was stated that, on comparing about 300 species of Touraine shells with about 450 from the Suffolk crag, 45 only were found to be common to both, which is in the proportion of only 15 per cent. The same small amount of agreement is found in the corals also. I formerly endeavoured to reconcile this marked difference in species with the supposed co-existence of the two faunas, by imagining them to have severally belonged to distinct zoological provinces or two seas, the one opening to the north, and the other to the south, with a barrier of land between them, like the Isthmus of Suez, separating the Red Sea and the Mediterranean. But I now abandon that idea for several reasons; among others, because I succeeded in 1841 in tracing the Crag fauna southwards in Normandy to within 70 miles of the Falunian type, near Dinan, yet found that both assemblages of fossils retained their distinctive characters, showing no signs of any blending of species or transition of climate.
On a comparison of 280 Mediterranean shells with 600 British species, made for me by an experienced conchologist in 1841, 160 were found to be common to both collections, which is in the proportion of 57 per cent., a fourfold greater specific resemblance than between the seas of the crag and the faluns, notwithstanding the greater geographical distance between England and the Mediterranean than between Suffolk and the Loire. The principal grounds, however, for referring the English crag to the older Pliocene and the French faluns to the Miocene epochs, consist in the predominance of fossil shells in the British strata identifiable with species, not only still living, but which are now inhabitants of neighbouring seas, while the accompanying extinct species are of genera such as characterize Europe. In the faluns, on the contrary, the recent species are in a decided minority, and many of them, like the associated extinct testacea, are much less European in character, and point to the prevalence of a warmer climate,—in other words, to a state of things receding farther from the present condition of Europe, geographically and climatologically, and doubtless, therefore, receding farther in time.
Bordeaux.—A great extent of country between the Pyrenees and the Gironde is overspread by tertiary deposits, which have been more particularly studied in the environs of Bordeaux and Dax, from whence about 700 species of shells have been obtained. A large proportion of these shells belong to the same zoological type as those of Touraine; but many are peculiar, and the whole may possibly constitute a somewhat older division of the Miocene period than the faluns of the Loire. We must wait, however, for farther investigations, in order to decide this question with accuracy.
Piedmont.—Many of the shells peculiar to the hill of the Superga, near Turin, agree with those found at Bordeaux and Dax; but the proportion of recent species is much less. The strata of the Superga consist of a bright green sand and marl, and a conglomerate with pebbles, chiefly of green serpentine, and are inclined at an angle of more than 70°. This formation, which attains a great thickness in the valley of the Bormida, is probably one of the oldest Miocene groups hitherto discovered.
Molasse of Switzerland.—If we cross the Alps, and pass from Piedmont to Savoy, we find there, at the northern base of the great chain, and throughout the lower country of Switzerland, a soft green sandstone much resembling some of the beds of the basin of the Bormida, above described, and associated in a similar manner with marls and conglomerate. This formation is called in Switzerland "molasse," said to be derived from "mol," "soft" because the stone is easily cut in the quarry. It is of vast thickness, and probably divisible into several formations. How large a portion of these belong to the Miocene period cannot yet be determined, as fossil shells are often entirely wanting. In some places a decided agreement of the fossil fishes of the molasse and faluns has been observed. Among those common to both, M. Agassiz pointed out to me Lamna contortidens, Myliobates Studeri, Spherodus cinctus, Notidanus primigenius, and others.
Lisbon.—Marine tertiary strata near Lisbon contain shells which agree very closely with those of Bordeaux, and are therefore referred to the Miocene era. Thus, out of 112 species collected by Mr. Smith of Jordanhill, between 60 and 70 were found to be common to the strata of Bordeaux and Dax, the recent species being in the proportion of 21 per cent.
Older Pliocene and Miocene formations in the United States.—Between the Alleghany mountains, formed of older rocks, and the Atlantic, there intervenes, in the United States, a low region occupied principally by beds of marl, clay, and sand, consisting of the cretaceous and tertiary formations, and chiefly of the latter. The general elevation of this plain bordering the Atlantic does not exceed 100 feet, although it is sometimes several hundred feet high. Its width in the middle and southern states is very commonly from 100 to 150 miles. It consists, in the South, as in Georgia, Alabama, and South Carolina, almost exclusively of Eocene deposits; but in North Carolina, Maryland, Virginia, and Delaware, more modern strata predominate, which I have assimilated in age to the English crag and Faluns of Touraine.[172-A] If, chronologically speaking, they can be truly said to be the representatives of these two European formations, they may range in age from the Older Pliocene to the Miocene epoch, according to the classification of European strata adopted in this chapter.
The proportion of fossil shells agreeing with recent, out of 147 species collected by me, amounted to about 17 per cent., or one-sixth of the whole; but as the fossils so assimilated were almost always the same as species now living in the neighbouring Atlantic, the number may hereafter be augmented, when the recent fauna of that ocean is better known. In different localities, also, the proportion of recent species varied considerably.