Of the singular family called Rudistes, by Lamarck, hereafter to be mentioned, as extremely characteristic of the chalk of Southern Europe, a single representative only ([fig. 213.]) has been discovered in the white chalk of England.

Hippurites Mortoni, Mantell. Houghton, Sussex. White chalk. Diameter one seventh of nat. size.

On the side where the shell is thinnest, there is one external furrow and corresponding internal ridge, a, b. [figs. 213], [214.]; but they are usually less prominent than in these figures. This species has been referred to Hippurites, but does not, I believe, fully agree in character with that genus. I have never seen the opercular piece, or valve, as it is called by those conchologists who regard the Rudistes as bivalve mollusca. The specimen above figured was discovered by the late Mr. Dixon.

The remains of fishes of the Upper Cretaceous formations consist chiefly of teeth of the shark family of genera, in part common to the tertiary, and partly distinct. But we meet with no bones of land animals, nor any terrestrial or fluviatile shells, nor any plants, except sea weeds, and here and there a piece of drift wood. All the appearances concur in leading us to conclude that the white chalk was the product of an open sea of considerable depth.

The existence of turtles and oviparous saurians, and of a Pterodactyl or winged-lizard, found in the white chalk of Maidstone, implies, no doubt, some neighbouring land; but a few small islets in mid-ocean, like Ascension, so much frequented by migratory droves of turtles, might perhaps have afforded the required retreat where these creatures might lay their eggs in the sand, or from which the flying species may have been blown out to sea. Of the vegetation of such islands we have scarcely any indication, but it consisted partly of cycadeous plants; for a fragment of one of these was found by Capt. Ibbetson in the chalk marl of the Isle of Wight, and is referred by A. Brongniart to Clathraria Lyellii, Mantell, a species common to the antecedent Wealden period.

Geographical extent and origin of the While Chalk.—The area over which the white chalk preserves a nearly homogeneous aspect is so vast, that the earlier geologists despaired of discovering any analogous deposits of recent date. Pure chalk, of nearly uniform aspect and composition, is met with in a north-west and south-east direction, from the north of Ireland to the Crimea, a distance of about 1140 geographical miles; and in an opposite direction it extends from the south of Sweden to the south of Bordeaux, a distance of about 840 geographical miles. In Southern Russia, according to Sir R. Murchison, it is sometimes 600 feet thick, and retains the same mineral character as in France and England, with the same fossils, including Inoceramus Cuvieri, Belemnites mucronatus, and Ostrea vesicularis.

But it would be an error to imagine, that the chalk was ever spread out continuously over the whole of the space comprised within these limits, although it prevailed in greater or less thickness over large portions of that area. On turning to those regions of the Pacific where coral reefs abound, we find some archipelagoes of lagoon islands, such as that of the Dangerous Archipelago, for instance, and that of Radack, with several adjoining groups, which are from 1100 to 1200 miles in length, and 300 or 400 miles broad; and the space to which Flinders proposed to give the name of the Corralline Sea is still larger; for it is bounded on the east by the Australian barrier—all formed of coral rock,—on the west by New Caledonia, and on the north by the reefs of Louisiade. Although the islands in these areas may be thinly sown, the mud of the decomposing zoophytes may be scattered far and wide by oceanic currents. That this mud would resemble chalk I have already hinted when speaking of the Faxoe limestone, [p. 211.]; and it was also remarked in an early part of this volume, that some even of that chalk which appears to an ordinary observer quite destitute of organic remains, is nevertheless, when seen under the microscope, full of fragments of corals and sponges; together with the valves of entomostraca, the shells of foraminifera, and still more minute infusoria.[215-A] (See [p. 26.])