The minor groups into which the Wealden has been commonly divided in England are, as before stated, three, and they succeed each other in the following descending order[227-A]:—
| Thickness. | ||
| 1st. | Weald Clay, sometimes including thin beds of sand and shelly limestone | 140 to 280 ft. |
| 2d. | Hastings Sand, in which occur some clays and calcareous grits | 400 to 500 ft. |
| 3d. | Purbeck Beds, consisting of various kinds of limestones and marls | 150 to 200 ft. |
Weald Clay.
The first division, or Weald Clay, is of purely freshwater origin. The uppermost beds are not only conformable, as Dr. Fitton observes, to the inferior strata of the Lower Greensand, but of similar mineral composition. To explain this, we may suppose, that as the delta of a great river was tranquilly subsiding, so as to allow the sea to encroach upon the space previously occupied by freshwater, the river still continued to carry down the same sediment into the sea. In confirmation of this view it may be stated, that the remains of the Iguanodon Mantelli, a gigantic terrestrial reptile, very characteristic of the Wealden, has been discovered near Maidstone, in the overlying Kentish rag, or marine limestone of the Lower Greensand. Hence we may infer that some of the saurians which inhabited the country of the great river continued to live when part of the country had become submerged beneath the sea. Thus, in our own times, we may suppose the bones of large alligators to be frequently entombed in recent freshwater strata in the delta of the Ganges. But if part of that delta should sink down so as to be covered by the sea, marine formations might begin to accumulate in the same space where freshwater beds had previously been formed; and yet the Ganges might still pour down its turbid waters in the same direction, and carry seaward the carcasses of the same species of alligator, in which case their bones might be included in marine as well as in subjacent freshwater strata.
The Iguanodon, first discovered by Dr. Mantell, has left more of its remains in the Wealden strata of the south-eastern counties, and Isle of Wight, than any other genus of associated saurians. It was an herbivorous reptile, and regarded by Cuvier as more extraordinary than any with which he was acquainted; for the teeth, though bearing a great analogy to the modern Iguanas which now frequent the tropical woods of America and the West Indies, exhibit many striking and important differences (see [fig. 230.]). It appears that they have been worn by mastication; whereas the existing herbivorous reptiles clip and gnaw off the vegetable productions on which they feed, but do not chew them. Their teeth, when worn, present an appearance of having been chipped off, and never, like the fossil teeth of the Iguanodon, have a flat ground surface (see [fig. 231.]), resembling the grinders of herbivorous mammalia. Dr. Mantell computes that the teeth and bones of this animal which have passed under his examination during the last twenty years, must have belonged to no less than seventy-one distinct individuals; varying in age and magnitude from the reptile just burst from the egg, to one of which the femur measured 24 inches in circumference. Yet notwithstanding that the teeth were more numerous than any other bones, it is remarkable that it was not till the relics of all these individuals had been found, that a solitary example of part of a jaw-bone was obtained. More recently remains both of the upper and lower jaw have been met with in the Hastings Beds in Tilgate Forest. Their size was somewhat greater than had been anticipated, and even allowing that the tail was short, which Professor Owen infers from the short bodies of the caudal vertebræ, Dr. Mantell estimates the probable length of some of these saurians at between 30 and 40 feet. The largest femur yet found measures 4 feet 8 inches in length, the circumference of the shaft being 25 inches, and round the condyles 42 inches.
Teeth of Iguanodon.
Fig. 230. Partially worn tooth of a young animal. (Mantell.)