Throughout North Wales the Wenlock shales rest unconformably upon the Caradoc sandstones; and the Caradoc is in its turn unconformable to the Llandeilo beds, showing a considerable interval of time between the deposition of this group and that of the formations next above and below it. The Caradoc sandstone in the neighbourhood of the Longmynd Hills in Shropshire, appears to Professor E. Forbes to have been a deep-sea deposit formed around the margin of high and steep land. That land consisted partly of upraised Llandeilo flags and partly of rocks of still older date.[359-B]
Such evidence of the successive disturbance of strata during the Silurian period in Great Britain is what we might look for when we have discovered the signs of so grand a series of volcanic eruptions as the contemporaneous greenstones and tuffs of the Welsh mountains afford.
Silurian Strata of the United States.
The position of some of these strata, where they are bent and highly inclined in the Appalachian chain, or where they are nearly horizontal to the west of that chain, is shown in the section, [fig. 379.] [p. 327.] But these formations can be studied still more advantageously north of the same line of section, in the states of New York, Ohio, and other regions north and south of the great Canadian lakes. Here they are found, as in Russia, in horizontal position, and are more rich in well-preserved fossils than in almost any spot in Europe. The American strata may readily be divided into Upper and Lower Silurian, corresponding in age and fossils to the European divisions bearing the same names. The subordinate members of the New York series, founded on lithological and geographical considerations, are most useful in the United States, but even there are only of local importance. Some few of them, however, tally very exactly with English divisions, as for example the limestone, over which the Niagara is precipitated at the great cataract, which, with its underlying shales, agrees paleontologically with the Wenlock limestone and shale of Siluria. There is also a marked general correspondence in the succession of fossil forms, and even species, as we trace the organic remains downwards from the highest to the lowest beds.
Mr. D. Sharpe, in his report on the mollusca collected by me from these strata in North America[359-C], has concluded that the number of species common to the Silurian rocks, on both sides of the Atlantic, is between 30 and 40 per cent.; a result which, although no doubt liable to future modification, when a larger comparison shall have been made, proves, nevertheless, that many of the species had a wide geographical range. It seems that comparatively few of the gasteropods and lamellibranchiate bivalves of North America can be identified specifically with European fossils, while no less than two-fifths of the brachiopoda are the same. In explanation of these facts, it is suggested, that most of the recent brachiopoda (especially the orthidiform ones) are inhabitants of deep water, and may have had a wider geographical range than shells living near shore. The predominance of bivalve mollusca of this peculiar class has caused the Silurian period to be sometimes styled the age of brachiopods.
Whether the Silurian rocks are of deep-water origin.—The grounds relied upon by Professor E. Forbes, for inferring that the larger part of the Silurian Fauna is indicative of a sea more than 70 fathoms deep, are the following: first, the small size of the greater number of conchifera; secondly, the paucity of pectinibranchiata (or spiral univalves); thirdly, the great number of floaters, such as Bellerophon, Orthoceras, &c.; fourthly, the abundance of orthidiform brachiopoda; fifthly, the absence or great rarity of fossil fish.
It is doubtless true that some living Terebratulæ, on the coast of Australia, inhabit shallow water; but all the known species, allied in form to the extinct Orthis, inhabit the depths of the sea. It should also be remarked that Mr. Forbes, in advocating these views, was well aware of the existence of shores, bounding the Silurian sea in Shropshire, and of the occurrence of littoral species of this early date in the northern hemisphere. Such facts are not inconsistent with his theory; for he has shown, in another work, how, on the coast of Lycia, deep-sea strata are at present forming in the Mediterranean, in the vicinity of high and steep land.
Had we discovered the ancient delta of some large Silurian river, we should doubtless have known more of the shallow, and brackish water, and fluviatile animals, and of the terrestrial flora of the period under consideration. To assume that there were no such deltas in the Silurian world, would be almost as gratuitous an hypothesis, as for the inhabitants of the coral islands of the Pacific to indulge in a similar generalization respecting the actual condition of the globe.[360-A]
Mineral Character of Silurian Strata.
In lithological character, the Silurian strata vary greatly when we trace them through Europe and North America. The shales called mudstones are as little altered from some deposits, found in recent submarine banks, as are those of many tertiary formations. We meet with red sandstone and red marl, with gypsum and salt, of Upper Silurian date, in the Niagara district, which might be mistaken for trias. The whitish granular sandstone at the base of the Silurian series in Sweden resembles the tertiary siliceous grit of Fontainebleau. The Calcareous Grit, oolite, and pisolite of Upper Silurian age in Gothland, are described by Sir R. Murchison as singularly like rocks of the oolitic period near Cheltenham; and, not to cite more examples, the Wenlock or Dudley limestone often resembles a modern coral-reef. If, therefore, uniformity of aspect has been thought characteristic of rocks of this age, the idea must have arisen from the similarity of feature acquired by strata subject to metamorphic action. This influence, seeing that the causes of change are always shifting the theatre of their principal development, must be multiplied throughout a wider geographical area by time, and become more general in any given system of rocks in proportion to their antiquity. We are now acquainted with dense groups of Eocene slates in the Alps, which were once mistaken by experienced geologists for Transition or Silurian formations. The error arose from attaching too great importance to mineral character as a test of age, for the tertiary slates in question having acquired that crystalline texture which is in reality most prevalent in the most ancient sedimentary formations.