CHAPTER XXXIV.
TRANSMUTATION OF SPECIES—Continued.
Recapitulation of the arguments in favor of the theory of transmutation of species—Their insufficiency—Causes of difficulty in discriminating species—Some varieties possibly more distinct than certain individuals of distinct species—Variability in a species consistent with a belief that the limits of deviation are fixed—No facts of transmutation authenticated—Varieties of the Dog—the Dog and Wolf distinct species—Mummies of various animals from Egypt identical in character with living individuals—Seeds and plants from the Egyptian tombs—Modifications produced in plants by agriculture and gardening.
The theory of the transmutation of species, considered in the last chapter, has met with some degree of favor from many naturalists, from their desire to dispense, as far as possible, with the repeated intervention of a First Cause, as often as geological monuments attest the successive appearance of new races of animals and plants, and the extinction of those pre-existing. But, independently of a predisposition to account, if possible, for a series of changes in the organic world by the regular action of secondary causes, we have seen that in truth many perplexing difficulties present themselves to one who attempts to establish the nature and reality of the specific character. And if once there appears ground of reasonable doubt, in regard to the constancy of species, the amount of transformation which they are capable of undergoing may seem to resolve itself into a mere question of the quantity of time assigned to the past duration of animate existence.
Before entering upon the reasons which may be adduced for rejecting Lamarck's hypothesis, I shall recapitulate, in a few words, the phenomena, and the whole train of thought, by which I conceive it to have been suggested, and which have gained for this and analogous theories, both in ancient and modern times, a considerable number of votaries.
In the first place, the various groups into which plants and animals may be thrown seem almost invariably, to a beginner, to be so natural, that he is usually convinced at first, as was Linnæus to the last, "that genera are as much founded in nature as the species which compose them."[800] When by examining the numerous intermediate gradations the student finds all lines of demarcation to be in most instances obliterated, even where they at first appeared most distinct, he grows more and more sceptical as to the real existence of genera, and finally regards them as mere arbitrary and artificial signs, invented, like those which serve to distinguish the heavenly constellations, for the convenience of classification, and having as little pretensions to reality.
Doubts are then engendered in his mind as to whether species may not also be equally unreal. The student is probably first struck with the phenomenon, that some individuals are made to deviate widely from the ordinary type by the force of peculiar circumstances, and with the still more extraordinary fact, that the newly acquired peculiarities are faithfully transmitted to the offspring. How far, he asks, may such variations extend in the course of indefinite periods of time, and during great vicissitudes in the physical condition of the globe? His growing incertitude is at first checked by the reflection that nature has forbidden the intermixture of the descendants of distinct original stocks, or has, at least, entailed sterility on their offspring, thereby preventing their being confounded together, and pointing out that a multitude of distinct types must have been created in the beginning, and must have remained pure and uncorrupted to this day.
Relying on this general law, he endeavors to solve each difficult problem by direct experiment, until he is again astounded by the phenomenon of a prolific hybrid, and still more by an example of a hybrid perpetuating itself throughout several generations in the vegetable world. He then feels himself reduced to the dilemma of choosing between two alternatives; either to reject the test, or to declare that the two species, from the union of which the fruitful progeny has sprung, were mere varieties. If he prefer the latter, he is compelled to question the reality of the distinctness of all other supposed species which differ no more than the parents of such prolific hybrids; for although he may not be enabled immediately to procure, in all such instances, a fruitful offspring; yet experiments show, that after repeated failures, the union of two recognized species may at last, under very favorable circumstances, give birth to a fertile progeny. Such circumstances, therefore, the naturalist may conceive to have occurred again and again, in the course of a great lapse of ages.