We have seen how many distinct geographical provinces there are of aquatic and terrestrial species, and how great are the powers of migration conferred on different classes, whereby the inhabitants of one region may be enabled from time to time to invade another, and do actually so migrate and diffuse themselves over new countries. Now, although our knowledge of the history of the animate creation dates from so recent a period, that we can scarcely trace the advance or decline of any animal or plant, except in those cases where the influence of man has intervened; yet we can easily conceive what must happen when some new colony of wild animals or plants enters a region for the first time, and succeeds in establishing itself.
Supposed effects of the first entrance of the polar bear into Iceland.—Let us consider how great are the devastations committed at certain periods by the Greenland bears, when they are drifted to the shores of Iceland in considerable numbers on the ice. These periodical invasions are formidable even to man; so that when the bears arrive, the inhabitants collect together, and go in pursuit of them with fire-arms—each native who slays one being rewarded by the King of Denmark. The Danes of old, when they landed in their marauding expeditions upon our coast, hardly excited more alarm, nor did our islanders muster more promptly for the defence of their lives and property against the common enemy, than the modern Icelanders against these formidable brutes. It often happens, says Henderson, that the natives are pursued by the bear when he has been long at sea, and when his natural ferocity has been heightened by the keenness of hunger; if unarmed, it is frequently by stratagem only that they make their escape.[965]
Let us cast our thoughts back to the period when the first polar bears reached Iceland, before it was colonized by the Norwegians in 874: we may imagine the breaking up of an immense barrier of ice like that which, in 1816 and the following year, disappeared from the east coast of Greenland, which it had surrounded for four centuries. By the aid of such means of transportation a great number of these quadrupeds might effect a landing at the same time, and the havoc which they would make among the species previously settled in the island would be terrific. The deer, foxes, seals, and even birds, on which these animals sometimes prey, would be soon thinned down.
But this would be a part only, and probably an insignificant portion, of the aggregate amount of change brought about by the new invader. The plants on which the deer fed, being less consumed in consequence of the lessened numbers of that herbivorous species, would soon supply more food to several insects, and probably to some terrestrial testacea, so that the latter would gain ground. The increase of these would furnish other insects and birds with food, so that the numbers of these last would be augmented. The diminution of the seals would afford a respite to some fish which they had persecuted; and these fish, in their turn, would then multiply and press upon their peculiar prey. Many water-fowls, the eggs and young of which are devoured by foxes, would increase when the foxes were thinned down by the bears; and the fish on which the water-fowls subsisted would then, in their turn, be less numerous. Thus the numerical proportions of a great number of the inhabitants, both of the land and sea, might be permanently altered by the settling of one new species in the region; and the changes caused indirectly would ramify through all classes of the living creation, and be almost endless.
An actual illustration of what we have here only proposed hypothetically, is in some degree afforded by the selection of small islands by the eider duck for its residence during the season of incubation, its nest being seldom if ever found on the shores of the main land, or even of a large island. The Icelanders are so well aware of this, that they have expended a great deal of labor in forming artificial islands, by separating from the main land certain promontories, joined to it by narrow isthmuses. This insular position is necessary to guard against the destruction of the eggs and young birds, by foxes, dogs, and other animals. One year, says Hooker, it happened that, in the small island of Vidoe, adjoining the coast of Iceland, a fox got over upon the ice, and caused great alarm, as an immense number of ducks were then sitting on their eggs or young ones. It was long before he was taken, which was at last, however, effected by bringing another fox to the island, and fastening it by a string near the haunt of the former, by which he was allured within shot of the hunter.[966]
The first appearance of a new species causes the chief disturbance.—It is usually the first appearance of an animal or plant, in a region to which it was previously a stranger, that gives rise to the chief alteration; since, after a time, an equilibrium is again established. But it must require ages before such a new adjustment of the relative forces of so many conflicting agents can be definitely settled. The causes in simultaneous action are so numerous, that they admit of an almost infinite number of combinations; and it is necessary that all these should have occurred once before the total amount of change, capable of flowing from any new disturbing force, can be estimated.
Thus, for example, suppose that once in two centuries a frost of unusual intensity, or a volcanic eruption of great violence accompanied by floods from the melting of glaciers, should occur in Iceland; or an epidemic disease, fatal to the larger number of individuals of some one species, and not affecting others,—these, and a variety of other contingencies, all of which may occur at once, or at periods separated by different intervals of time, ought to happen before it would be possible for us to declare what ultimate alteration the presence of any new comer, such as the bear before mentioned, might occasion in the animal population of the isle.
Every new condition in the state of the organic or inorganic creation, a new animal or plant, an additional snow-clad mountain, any permanent change, however slight in comparison to the whole, gives rise to a new order of things, and may make a material change in regard to some one or more species. Yet a swarm of locusts, or a frost of extreme intensity, or an epidemic disease, may pass away without any great apparent derangement; no species may be lost, and all may soon recover their former relative numbers, because the same scourges may have visited the region again and again, at preceding periods. Every plant that was incapable of resisting such a degree of cold, every animal which was exposed to be entirely cut off by an epidemic or by famine caused by the consumption of vegetation by the locusts, may have perished already, so that the subsequent recurrence of similar catastrophes is attended only by a temporary change.
Changes caused by Man
We are best acquainted with the mutations brought about by the progress of human population, and the growth of plants and animals favored by man. To these, therefore, we should in the first instance turn our attention. If we conclude, from the concurrent testimony of history and of the evidence yielded by geological data, that man is, comparatively speaking, of very modern origin, we must at once perceive how great a revolution in the state of the animate world the increase of the human race, considered merely as consumers of a certain quantity of organic matter, must necessarily cause.