FORMATION OF CORAL REEFS.

Growth of coral chiefly confined to tropical regions—Principal genera of coral-building zoophytes—Their rate of growth—Seldom flourish at greater depths than twenty fathoms—Atolls or annular reefs with lagoons—Maldive Isles—Origin of the circular form—Coral reefs not based on submerged volcanic craters—Mr. Darwin's theory of subsidence in explanation of atolls, encircling and barrier reefs—Why the windward side of atolls highest—Subsidence explains why all atolls are nearly on one level—Alternate areas of elevation and subsidence—Origin of openings into the lagoons—Size of atolls and barrier reefs—Objection to the theory of subsidence considered—Composition, structure, and stratified arrangement of rocks now forming in coral reefs—Lime, whence derived—Supposed increase of calcareous matter in modern epochs controverted—Concluding remarks.

The powers of the organic creation in modifying the form and structure of the earth's crust, are most conspicuously displayed in the labors of the coral animals. We may compare the operation of these zoophytes in the ocean, to the effects produced on a smaller scale upon the land by the plants which generate peat. In the case of the Sphagnum, the upper part vegetates while the lower part is entering into a mineral mass, in which the traces of organization remain when life has entirely ceased. In corals, in like manner, the more durable materials of the generation that has passed away serve as the foundation on which the living animals continue to rear a similar structure.

The stony part of the lamelliform zoophyte may be likened to an internal skeleton; for it is always more or less surrounded by a soft animal substance capable of expanding itself; yet, when alarmed, it has the power of contracting and drawing itself almost entirely into the cells and hollows of the hard coral. Although oftentimes beautifully colored in their own element, the soft parts become when taken from the sea nothing more in appearance than a brown slime spread over the stony nucleus.[1111]

The growth of those corals which form reefs of solid stone is entirely confined to the warmer regions of the globe, rarely extending beyond the tropics above two or three degrees, except under peculiar circumstances, as in the Bermuda Islands, in lat. 32° N., where the Atlantic is warmed by the Gulf stream. The Pacific Ocean, throughout a space comprehended between the thirtieth parallels of latitude on each side of the equator, is extremely productive of coral; as also are the Arabian and Persian Gulfs. Coral is also abundant in the sea between the coast of Malabar and the island of Madagascar. Flinders describes a reef of coral on the east coast of New Holland as having a length of nearly 1000 miles, and as being in one part unbroken for a distance of 350 miles. Some groups of coral islands in the Pacific are from 1100 to 1200 miles Fig. 108.

Meandrina labyrinthica, Lam. in length, by 300 or 400 in breadth, as the Dangerous Archipelago, for example, and that called Radack by Kotzebue; but the islands within these spaces are always small points, and often very thinly sown.

Of the numerous species of zoophytes which are engaged in the production of coral banks, some of the most common belong to the Lamarckian genera Astrea, Porites, Madrepora, Millepora, Caryophyllia, and Meandrina.

Rate of the growth of Coral.—Very different opinions have been entertained in regard to the rate at which coral reefs increase. In Captain Beechey's late expedition to the Pacific, no positive information could be obtained of any channel having been filled up within a given period; and it seems established, that several reefs had remained for more than half a century, at about the same depth from the surface.

Ehrenberg also questions the fact of channels and harbors having been closed up in the Red Sea by the rapid increase of coral limestone. He supposes the notion to have arisen from the circumstance of havens having been occasionally filled up in some places with coral sand, in others with large quantities of ballast of coral rock thrown down from vessels.