A still more anomalous state of things presents itself in the southern hemisphere. Even in the temperate zone, between the latitudes 52° and 56° S., as, for example, in Tierra del Fuego, as well as in the woody region immediately north of the Straits of Magellan, and in the Falkland Islands, no reptiles of any kind are met with, not even a snake, lizard, or frog; but in these same countries we find the guanaco (a kind of llama), a deer, the puma, a large species of fox, many small rodentia, besides the seal and otter, together with the porpoise, whale, and other cetacea.
On what grand laws in the animal physiology these remarkable phenomena depend, cannot in the present state of science be conjectured; nor could we predict whether any opposite condition of the atmosphere, in respect to heat, moisture, and other circumstances, would bring about a state of animal life which might be called the converse of that above described, namely, a state in which reptiles of every size and order might abound, and mammalia disappear.
The nearest approximation to such a fauna is found in the Galapagos Archipelago. These islands, situated under the equator, and nearly 600 miles west of the coast of Peru, have been called "the land of reptiles," so great is the number of snakes, large tortoises, and lizards, which they support. Among the lizards, the first living species proper to the ocean has been discovered. Yet, although some of these islands are from 3000 to 4000 feet high, and one of them 75 miles long, they contain, with the exception of one small mouse, no indigenous mammifer. Even here, however, it is true that in the neighboring sea there are seals, and several kinds of cetacea.[221]
It may be unreasonable to look for a nearer analogy between the fauna now existing in any part of the globe, and that which we can show to have prevailed when our secondary strata were deposited, because we must always recollect that a climate like that now experienced at the equator, coexisting with the unequal days and nights of European latitudes, was a state of things to which there is now no counterpart on the globe. Consequently, the type of animal and vegetable existence required for such a climate might be expected to deviate almost as widely from that now established, as do the flora and fauna of our tropical differ from those of our arctic regions.
In the Tertiary strata.—The tertiary formations were deposited when the physical geography of the northern hemisphere had been entirely altered. Large inland lakes had become numerous, as in central France and other countries. There were gulfs of the sea, into which considerable rivers emptied themselves, and where strata like those of the Paris basin were accumulated. There were also formations in progress, in shallow seas not far from shore, such as are indicated by portions of the Faluns of the Loire, and the English Crag.
The proximity, therefore, of large tracts of dry land to the seas and lakes then existing, may, in a great measure, explain why the remains of land animals, so rare in the older strata, are not uncommon in these more modern deposits. Yet even these have sometimes proved entirely destitute of mammiferous relics for years after they had become celebrated for the abundance of their fossil testacea, fish, and reptiles. Thus the calcaire grossier, a marine limestone of the district round Paris, had afforded to collectors more than 1100 species of shells, besides many zoophytes, echinodermata, and the teeth of fish, before the bones of one or two land quadrupeds were met with in the same rock. The strata called London and Plastic clay in England have been studied for more than half a century, and about 400 species of shells, 50 or more of fish, besides several kinds of chelonian and saurian reptiles, were known before a single mammifer was detected. At length, in the year 1839, there were found in this formation the remains of a monkey, an opossum, a bat,[222] and a species of the extinct genus Hyracotherium, allied to the Peccary or hog tribe.
If we examine the strata above the London clay in England, we first meet with mammiferous remains in the Isle of Wight, in beds also belonging to the Eocene epoch, such as the remains of the Palæotherium, Anoplotherium, and other extinct quadrupeds, agreeing very closely with those first found by Cuvier, near Paris, in strata of the same age, and of similar freshwater origin.
In France we meet with another fauna, both conchological and mammalian in the Miocene "faluns" of the Loire; above which in the ascending series in Great Britain we arrive at the coralline crag of Suffolk, a marine formation which has yielded three or four hundred species of shells, very different from the Eocene testacea, and of which a large proportion, although a minority of the whole number, are recent, besides many corals, echini, foraminifera, and fish, but as yet no relic decidedly mammalian except the ear-bone of a whale.
In the shelly sand, provincially termed "Red Crag," in Suffolk, which immediately succeeds the coralline, constituting a newer member of the same tertiary group, about 250 species of shells have been recognized, of which a still larger proportion are recent. They are associated with numerous teeth of fish; but no signs of a warm-blooded quadruped had been detected until 1839, when the teeth of a leopard, a bear, a hog, and a species of ruminant, were found at Newbourn, in Suffolk, and since that time, several other genera of mammalia have been met with in the same formation, or in the Red Crag.[223]
Of a still newer date is the Norwich Crag, a fluvio-marine deposit of the Pleiocene epoch, containing a mixture of marine, fluviatile, and land shells, of which 90 per cent. or more are recent. These beds, since the time of their first investigation, have yielded a supply of mammalian bones of the genera mastodon, elephant, rhinoceros, pig, horse, deer, ox, and others, the bodies of which may have been washed down into the sea by rivers draining land, of which the contiguity is indicated by the occasional presence of terrestrial and freshwater shells.