In a memoir by Professor E. Forbes, before cited, he observes, that the land of passage by which the plants and animals migrated into Ireland consisted of the upraised marine drift which had previously formed the bottom of the glacial sea. Portions of this drift extend to the eastern shores of Wicklow and Wexford, others are found in the Isle of Man full of arctic shells, others on the British coast opposite Ireland. The freshwater marl, containing numerous skeletons of the great deer, or Megaceros, overlie in the Isle of Man that marine glacial drift. Professor Forbes also remarks that the subsequent disjunction of Ireland from England, or the formation of the Irish Channel, which is less than 400 feet in its greatest depth, preceded the opening of the Straits of Dover, or the final separation of England from the Continent. This he inferred from the present distribution of species both in the animal and vegetable kingdoms. Thus, for example, there are twice as many reptiles in Belgium as in England, and the number inhabiting England is twice that found in Ireland. Yet the Irish species are all common to England, and all the English to Belgium. It is therefore assumed that the migration of species westward having been the work of time, there was not sufficient lapse of ages to complete the fusion of the continental and British reptilian fauna, before France was separated from England and England from Ireland.

For the same reason there are also a great number of birds of short flight, and small quadrupeds, inhabiting England which do not cross to Ireland, the Irish Channel seeming to have arrested them in their westward course.*

(* E. Forbes, Fauna and Flora of British Isles, "Memoir of
the Geological Survey" volume 1 1846 page 344.)

The depth of the Irish Channel in the narrower parts is only 360 feet, and the English Channel between Dover and Calais less than 200, and rarely anywhere more than 300 feet; so that vertical movements of slight amount compared to some of those previously considered, with the aid of denuding operations or the waste of sea cliffs, and the scouring out of the channel, might in time effect the insulation of the lands above alluded to.

TIME REQUIRED FOR SUCCESSIVE CHANGES IN PHYSICAL GEOGRAPHY IN THE PLEISTOCENE PERIOD.

The time which it would require to bring about such changes of level, according to the average rate assumed in Chapter 3, however vast, will not be found to exceed that which would best explain the successive fluctuations in terrestrial temperature, the glaciation of solid rocks, the transportation of erratics above and below the sea-level, the height of arctic shells above the sea, and last, not least, the migration of the existing species of animals and plants into their actual stations, and the extinction of some conspicuous forms which flourished during the Pleistocene ages. When we duly consider all these changes which have taken place since the beginning of the glacial epoch, or since the forest of Cromer and the Elephas meridionalis flourished, we shall find that the phenomena become more and more intelligible in proportion to the slowness of the rate of elevation and depression which we assume.

The submergence of Wales to the extent of 1400 feet, as proved by glacial shells, would require 56,000 years, at the rate of 2 1/2 feet per century; but taking Professor Ramsay's estimate of 800 feet more, that depression being implied by the position of some of the stratified drift, we must demand an additional period of 32,000 years, amounting in all to 88,000; and the same time would be required for the re-elevation of the tract to its present height. But if the land rose in the second continental period as much as 600 feet above its present level, as in Figure 41, this 600 feet, first of rising and then of sinking, would require 48,000 years more; the whole of the grand oscillation, comprising the submergence and re-emergence, having taken about 224,000 years for its completion; and this, even if there were no pause or stationary period, when the downward movement ceased, and before it was converted into an upward one.

I am aware that it may be objected that the average rate here proposed is a purely arbitrary and conjectural one, because, at the North Cape, it is supposed that there has been a rise of about 5 feet in a century, and at Spitsbergen, according to Mr. Lamont, a still faster upheaval during the last 400 years.*

(* "Seasons with the Sea-Horses" page 202.)

But, granting that in these and some exceptional cases (none of them as yet very well established) the rising or sinking has, for a time, been accelerated, I do not believe the average rate of motion to exceed that above proposed. Mr. Darwin, I find, considers that such a mean rate of upheaval would be as high as we could assume for the west coast of South America, where we have more evidence of sudden changes of level than anywhere else. He has not, however, attempted to estimate the probable rate of secular elevation in that or any other region.