To this objection Lamarck would, no doubt, have answered that there had not been time for bringing about so great an amount of variation; for when Cuvier and some other of his contemporaries appealed to the embalmed animals and plants taken from Egyptian tombs, some of them 3000 years old, which had not experienced in that long period the slightest modification in their specific characters, he replied that the climate and soil of the valley of the Nile had not varied in the interval, and that there was therefore no reason for expecting that we should be able to detect any change in the fauna and flora. "But if," he went on to say, "the physical geography, temperature, and other conditions of life had been altered in Egypt as much as we know from geology has happened in other regions, some of the same animals and plants would have deviated so far from their pristine types as to be thought entitled to take rank as new and distinct species."
Although I cited this answer of Lamarck in my account of his theory,* I did not at the time fully appreciate the deep conviction which it displays of the slow manner in which geological changes have taken place and the insignificance of thirty or forty centuries in the history of a species, and that, too, at a period when very narrow views were entertained of the extent of past time by most of the ablest geologists, and when great revolutions of the earth's crust, and its inhabitants, were generally attributed to sudden and violent catastrophes.
(* Ibid. page 587.)
While in 1832 I argued against Lamarck's doctrine of the gradual transmutation of one species into another, I agreed with him in believing that the system of changes now in progress in the organic world would afford, when fully understood, a complete key to the interpretation of all the vicissitudes of the living creation in past ages. I contended against the doctrine, then very popular, of the sudden destruction of vast multitudes of species and the abrupt ushering into the world of new batches of plants and animals.
I endeavoured to sketch out (and it was, I believe, the first systematic attempt to accomplish such a task) the laws which govern the extinction of species, with a view of showing that the slow but ceaseless variations now in progress in physical geography, together with the migration of plants and animals into new regions, must in the course of ages give rise to the occasional loss of some of them and eventually cause an entire fauna and flora to die out; also that we must infer from geological data that the places thus left vacant from time to time are filled up without delay by new forms adapted to new conditions, sometimes by immigration from adjoining provinces, sometimes by new creations. Among the many causes of extinction enumerated by me were the power of hostile species, diminution of food, mutations in climate, the conversion of land into sea and of sea into land, etc. I firmly opposed Brocchi's hypothesis of a decline in the vital energy of each species;* maintaining that there was every reason to believe that the reproductive powers of the last surviving representatives of a species were as vigorous as those of their predecessors, and that they were as capable, under favourable circumstances, of repeopling the earth with their kind.
(* "Principles of Geology" 1st edition volume 2 chapter 8;
and 9th edition page 668.)
The manner in which some species are now becoming scarce and dying out, one after the other, appeared to me to favour the doctrine of the fixity of the specific character, showing a want of pliancy and capability of varying, which ensured their annihilation whenever changes adverse to their well-being occurred; time not being allowed for such a transformation as might be conceived capable of adapting them to the new circumstances, and of converting them into what naturalists would call new species.*
(* Laws of Extinction, "Principles of Geology" 1st edition
1832 volume 2 chapters 5 to 11 inclusive; and 9th edition
chapters 37 to 42 inclusive 1853.)
But while rejecting transmutation, I was equally opposed to the popular theory that the creative power had diminished in energy, or that it had been in abeyance ever since Man had entered upon the scene. That a renovating force which had been in full operation for millions of years should cease to act while the causes of extinction were still in full activity, or even intensified by the accession of Man's destroying power, seemed to me in the highest degree improbable. The only point on which I doubted was whether the force might not be intermittent instead of being, as Lamarck supposed, in ceaseless operation. Might not the births of new species, like the deaths of old ones, be sudden? Might they not still escape our observation? If the coming in of one new species, and the loss of one other which had endured for ages, should take place annually, still, assuming that there are a million of animals and plants living on the globe, it would require, I observed, a million of years to bring about a complete revolution in the fauna and flora. In that case, I imagined that, although the first appearance of a new form might be as abrupt as the disappearance of an old one, yet naturalists might never yet have witnessed the first entrance on the stage of a large and conspicuous animal or plant, and as to the smaller kinds, many of them may be conceived to have stolen in unseen, and to have spread gradually over a wide area, like species migrating into new provinces.*
(* "Principles of Geology" 1st edition 1832 volume 2 chapter
11; and 9th edition page 706.)