The Antiquity of Man

by Sir Charles Lyell

It had usually been supposed by the advocates of the immutability of species that domesticated races, if allowed to run wild, always revert to their parent type. Mr. Wallace had said in reply that a domesticated species, if it loses the protection of Man, can only stand its ground in a wild state by resuming those habits and recovering those attributes which it may have lost when under domestication. If these faculties are so much enfeebled as to be irrecoverable it will perish; if not and if it can adapt itself to the surrounding conditions, it will revert to the state in which Man first found it: for in one, two, or three thousand years, which may have elapsed since it was originally tamed, there will not have been time for such geographical, climatal, and organic changes as would only be suited to a new race or a new and allied species.

But in regard to plants Dr. Hooker questions the fact of reversion. According to him, species in general do not readily vary, but when they once begin to do so the new varieties, as every horticulturist knows, show a great inclination to go on departing more and more from the old stock. As the best marked varieties of a wild species occur on the confines of the area which it inhabits, so the best marked varieties of a cultivated plant are those last produced by the gardener. Cabbages, for example, wall fruits, and cereal, show no disposition, when neglected, to assume the characters of the wild states of these plants. Hence the difficulty of determining what are the true parent species of most of our cultivated plants. Thus the finer kinds of apples, if grown from seed, degenerate and become crabs, but in so doing they do not revert to the original wild crab-apple, but become crab states of the varieties to which they belong.*

(* "Introductory Essay to the Flora of Australia" page 9.)

It would lead me into too long a digression were I to attempt to give a fuller analysis of this admirable essay; but I may add that none of the observations are more in point, as bearing on the doctrine of what Hooker terms "creation by variation," than the great extent to which the internal characters and properties of plants, or their physiological constitution, are capable of being modified, while they exhibit externally no visible departure from the normal form. Thus, in one region a species may possess peculiar medicinal qualities which it wants in another, or it may be hardier and better able to resist cold. The average range in altitude, says Hooker, of each species of flowering plant in the Himalayan Mountains, whether in the tropical, temperate, or Alpine region, is 4000 feet, which is equivalent to twelve degrees of isothermals of latitude. If an individual of any of these species be taken from the upper limits of its range and carried to England, it is found to be better able to stand our climate than those from the lower or warmer stations. When several of these internal or physiological modifications are accompanied by variation in size, habits of growth, colour of the flowers, and other external characters, and these are found to be constant in successive generations, botanists may well begin to differ in opinion as to whether they ought to regard them as distinct species or not.

ALTERNATION OF GENERATIONS.

Hitherto, no rival hypothesis has been proposed as a substitute for the doctrine of transmutation; for what we term "independent creation," or the direct intervention of the Supreme Cause, must simply be considered as an avowal that we deem the question to lie beyond the domain of science.

The discovery by Steenstrup of alternate generation enlarges our views of the range of metamorphosis through which a species may pass, so that some of its stages (as when a Sertularia and a Medusa interchange) deviate so far from others as to have been referred by able zoologists to distinct genera, or even families. But in all these cases the organism, after running through a certain cycle of change, returns to the exact point from which it set out, and no new form or species is thereby introduced into the world. The only secondary cause therefore which has as yet been even conjecturally brought forward, to explain how in the ordinary course of nature a new specific form may be generated is, as Lamarck declared, "variation," and this has been rendered a far more probable hypothesis by the way in which "natural selection" is shown to give intensity and permanency to certain varieties.

INDEPENDENT CREATION.

When I formerly advocated the doctrine that species were primordial creations and not derivative, I endeavoured to explain the manner of their geographical distribution, and the affinity of living forms to the fossil types nearest akin to them in the Tertiary strata of the same part of the globe, by supposing that the creative power, which originally adapts certain types to aquatic and others to terrestrial conditions, has at successive geological epochs introduced new forms best suited to each area and climate, so as to fill the places of those which may have died out.

In that case, although the new species would differ from the old (for these would not be revived, having been already proved by the fact of their extinction to be incapable of holding their ground), still they would resemble their predecessors generically. For, as Mr. Darwin states in regard to new races, those of a dominant type inherit the advantages which made their parent species flourish in the same country, and they likewise partake in those general advantages which made the genus to which the parent species belonged a large genus in its own country.