Insects in European Coal.—Articulate animals of the genus Scorpion were found by Count Sternberg in 1835 in the coal-measures of Bohemia, and about the same time in those of Coalbrook Dale by Mr. Prestwich, were also true insects, such as beetles of the family Curculionidæ, a neuropterous insect of the genus Corydalis, and another related to the Phasmidæ, have been found.
From the coal of Wetting, in Westphalia, several specimens of the cockroach or Blatta family, and the wing of a cricket (Acridites) have been described by Germar. Professor Goldenberg published, in 1854, descriptions of no less than twelve species of insects from the nodular clay-ironstone of Saarbrück, near Trèves.[[4]] Among them are several Blattinæ, three species of Neuroptera, one beetle of the Scarabæus family, a grasshopper or locust, Gryllacris (see Fig. 435), and several white ants or Termites. Professor Goldenberg showed me, in 1864, the wing of a white ant, found low down in the productive coal-measures of Saarbrück, in the interior of a flattened Lepidodendron. It is much larger than that of any known living species of the same genus.
Batrachian Reptiles in Coal.—No vertebrated animals more highly organised than fish were known in rocks of higher antiquity than the Permian until the year 1844, when the Apateon pedestris, Meyer, was discovered in the coal-measures of Munster-Appel in Rhenish Bavaria, and three years later, in 1847, Professor von Dechen found three other distinct species of the same family of Amphibia in the Saarbruck coal-field above alluded to. These were described by the late Professor Goldfuss under the generic name of Archegosaurus. The skulls, teeth, and the greater portions of the skeleton, nay, even a large part of the skin, of two of these reptiles have been faithfully preserved in the centre of spheroidal concretions of clay-ironstone. The largest of these, Archegosaurus Decheni, must have been three feet six inches long. Figure 436 represents the skull and neck bones of the smallest of the three, of the natural size. They were considered by Goldfuss as saurians, but by Herman von Meyer as most nearly allied to the Labyrinthodon before mentioned ([p. 371]), and the remains of the extremities leave no doubt they were quadrupeds, “provided,” says Von Meyer, “with hands and feet terminating in distinct toes; but these limbs were weak, serving only for swimming or creeping.” The same anatomist has pointed out certain points of analogy between their bones and those of the Proteus anguinus; and Professor Owen has observed that they make an approach to the Proteus in the shortness of their ribs. Two specimens of these ancient reptiles retain a large part of the outer skin, which consisted of long, narrow, wedge-shaped, tile-like, and horny scales, arranged in rows (see Fig. 437).
In 1865, several species belonging to three different genera of the same family of perennibranchiate Batrachians were found in the coal-field of Kilkenny in bituminous shale at the junction of the coal with the underlying Stigmaria-bearing clay. They were, probably, inhabitants of a marsh, and the large processes projecting from the vertebræ of their tail imply, according to Professor Huxley, great powers of swimming. They were of the Labyrinthodont family, and their association with the fish of the coal, of which so large a proportion are ganoids, reminds us that the living perennibranchiate amphibia of America frequent the same rivers as the ganoid Lepidostei or bony pikes.
Labyrinthodont footprints in coal-measures.—In 1844, the very year when the Apateon, before mentioned, of the coal was first met with in the country between the Moselle and the Rhine, Dr. King published an account of the footprints of a large reptile discovered by him in North America. These occur in the coal-strata of Greensburg, in Westmoreland County, Pennsylvania; and I had an opportunity of examining them when in that country in 1846. The footmarks were first observed standing out in relief from the lower surface of slabs of sandstone, resting on thin layers of fine unctuous clay. I brought away one of these masses, which is represented in Fig. 438. It displays, together with footprints, the casts of cracks (a, a′) of various sizes. The origin of such cracks in clay, and casts of the same, has before been explained, and referred to the drying and shrinking of mud, and the subsequent pouring of sand into open crevices. It will be seen that some of the cracks, as at b, c, traverse the footprints, and produce distortion in them, as might have been expected, for the mud must have been soft when the animal walked over it and left the impressions; whereas, when it afterwards dried up and shrank, it would be too hard to receive such indentations.
