In regard to the two types, the Old Red Sandstone and the Devonian, I shall first treat of them separately, and then allude to the proofs of their having been to a great extent contemporaneous. That they constitute a series of rocks intermediate in date between the lowest Carboniferous and the uppermost Silurian is not disputed by the ablest geologists; and it can no longer be contended that the Upper, Middle, and Lower Old Red Sandstone preceded in date the three divisions to which, by aid of the marine shells, the Devonian rocks have been referred, while, on the other hand, we have not yet data for enabling us to affirm to what extent the subdivisions of the one series may be the equivalents in time of those of the other.

Upper Old Red Sandstone.—The highest beds of the series in Scotland, lying immediately below the coal in Fife, are composed of yellow sandstone well seen at Dura Den, near Coupar, in Fife, where, although the strata contain no mollusca, fish have been found abundantly, and have been referred to the genera Holoptychius, Pamphractus, Glyptopomus, and many others. In the county of Cork, in Ireland, a similar yellow sandstone occurs containing fish of genera characteristic of the Scotch Old Red Sandstone, as for example Coccosteus (a form represented by many species in the Old Red Sandstone and by one only in the Carboniferous group), and Glytolepis and Asterolepis, both exclusively confined to the “Old Red.” In the same Irish sandstone at Kiltorkan has been found an Anodonta or fresh-water mussel, the only shell hitherto discovered in the Old Red Sandstone of the British Isles (see Fig. 494).

In the same formation are found the fern (Fig. 496) and the Lepidodendron (Fig. 495), and other species of plants, some of which, Professor Heer remarks, agree specifically with species from the lower carboniferous beds. This induces him to lean to the opinion long ago advocated by Sir Richard Griffiths, that the yellow sandstone, in spite of its fish remains, should be classed as Lower Carboniferous, an opinion which I am not yet prepared to adopt. Between the Mountain Limestone and the yellow sandstone in the south-west of Ireland there intervenes a formation no less than 5000 feet thick, called the “Carboniferous slate,” and at the base of this, in some places, are local deposits, such as the Glengariff Grits, which appear to be beds of passage between the Carboniferous and Old Red Sandstone groups.

It is a remarkable result of the recent examination of the fossil flora of Bear Island, latitude 74° 30′ N., that Professor Heer has described as occurring in that part of the Arctic region (nearly twenty-six degrees to the north of the Irish locality) a flora agreeing in several of its species with that of the yellow sandstones of Ireland. This Bear Island flora is believed by Professor Heer to comprise species of plants some of which ascend even to the higher stages of the European Carboniferous formation, or as high as the Mountain Limestone and Millstone Grit. Palæontologists have long maintained that the same species which have a wide range in space are also the most persistent in time, which may prepare us to find that some plants having a vast geographical range may also have endured from the period of the Upper Devonian to that of the Millstone Grit.

Outliers of the Upper “Old Red” occur unconformably on older members of the group, and the formation represented at Whiteness, near Arbroath, a, [Fig. 55,] may probably be one of these outliers, though the want of organic remains renders this uncertain. It is not improbable that the beds given in this section as Nos. 1, 2, and 3, may all belong to the early part of the period of the Upper Old Red, as some scales of Holoptychius nobilissimus have been found scattered through these beds, No. 2, in Strathmore. Another nearly allied Holoptychius occurs in Dura Den, see Fig. 498 of this fish and also Fig. 497 of one of its scales, as these last are often the only parts met with; being scattered in Forfarshire through red-coloured shales and sandstones, as are scales of a large species of the same genus in a corresponding matrix in Herefordshire.[[1]] The number of fish obtained from the British Upper Old Red Sandstone amounts to fifteen species referred to eleven genera.