Among the peculiar lamellibranchiate bivalves common to the Plymouth limestone of Devonshire and the Continent, we find the Megalodon (Fig. 517). There are also twelve genera of Gasteropods which have yielded 36 species, four of which pass to the Carboniferous group, namely Macrocheilus, Acroculia, Euomphalus, and Murchisonia. Pteropods occur, such as Conularia (Fig. 518), and Cephalopods, such as Cyrtoceras, Gyroceras, Orthoceras, and others, nearly all of genera distinct from those prevailing in the Upper Devonian Limestone, or Clymenien-kalk of the Germans already mentioned. Although but few species of Trilobites occur, the characteristic Bronteus flabellifer (Fig. 519) is far from rare, and all collectors are familiar with its fan-like tail. In this same group, called, as before stated, the Stringocephalus, or Eifel Limestone, in Germany, several fish remains have been detected, and among others the remarkable genus Coccosteus, covered with its tuberculated bony armour; and these ichthyolites serve, as Sir R. Murchison observes (Siluria, p. 362), to identify this middle marine Devonian with the Old Red Sandstone of Britain and Russia.

Beneath the Eifel Limestone (the great central and typical member of “the Devonian” on the Continent) lie certain schists called by German writers “Calceola-schiefer,” because they contain in abundance a fossil body of very curious structure, Calceola sandalina (Fig. 520), which has been usually considered a brachiopod, but which some naturalists have lately referred to a Goniophyllum, supposing it to be an abnormal form of the order Zoantharia rugosa (see [Fig. 474]), differing from all other corals in being furnished with a strong operculum. This is by no means a rare fossil in the slaty limestone of South Devon, and, like the Eifel form, is confined to the middle group of this country.

Lower Devonian Rocks.—A great series of sandstones and glossy slates, with Crinoids, Brachiopods, and some corals, occurring on the coast at Lynmouth and the neighbourhood, and called the Lynton Group (see Table [p. 449], form the lowest member of the Devonian in North Devon. Among the 18 species of all classes enumerated by Mr. Etheridge, two-thirds are common to the Middle Devonian, but only one, the ubiquitous Atrypa reticularis, can with certainty be identified with Silurian species. Among the characteristic forms are Alveolites suborbicularis, also common to this formation in the Rhine, and Orthis arcuata, very widely spread in the North Devon localities. But we may expect a large addition to the number of fossils whenever these strata shall have been carefully searched. The Spirifer Sandstone of Sandberger, as exhibited in the rocks bordering the Rhine between Coblentz and Caub, belong to this Lower division, and the same broad-winged Spirifers distinguish the Devonian strata of North America.

Among the Trilobites of this era several large species of Homalonotus (Fig. 522) are conspicuous. The genus is still better known as a Silurian form, but the spinose species appear to belong exclusively to the “Lower Devonian,” and are found in Britain, Europe, and the Cape of Good Hope.

Devonian of Russia.—The Devonian strata of Russia extend, according to Sir R. Murchison, over a region more spacious than the British Isles; and it is remarkable that, where they consist of sandstone like the “Old Red” of Scotland and Central England, they are tenanted by fossil fishes often of the same species and still oftener of the same genera as the British, whereas when they consist of limestone they contain shells similar to those of Devonshire, thus confirming, as Sir Roderick has pointed out, the contemporaneous origin which had been previously assigned to formations exhibiting two very distinct mineral types in different parts of Britain.[[6]] The calcareous and the arenaceous rocks of Russia above alluded to alternate in such a manner as to leave no doubt of their having been deposited in different parts of the same great period.