Another important discovery was soon afterwards made by Brocchi in Italy, who investigated the argillaceous and sandy deposits, replete with shells, which form a low range of hills, flanking the Apennines on both sides, from the plains of the Po to Calabria. These lower hills were called by him the Subapennines, and were formed of strata chiefly marine, and newer than those of Paris and London.
Another tertiary group occurring in the neighbourhood of Bordeaux and Dax, in the South of France, was examined by M. de Basterot in 1825, who described and figured several hundred species of shells, which differed for the most part both from the Parisian series and those of the Subapennine hills. It was soon, therefore, suspected that this fauna might belong to a period intermediate between that of the Parisian and Subapennine strata, and it was not long before the evidence of superposition was brought to bear in support of this opinion; for other strata, contemporaneous with those of Bordeaux, were observed in one district (the Valley of the Loire), to overlie the Parisian formation, and in another (in Piedmont) to underlie the Subapennine beds. The first example of these was pointed out in 1829 by M. Desnoyers, who ascertained that the sand and marl of marine origin called faluns, near Tours, in the basin of the Loire, full of sea-shells and corals, rested upon a lacustrine formation, which constitutes the uppermost subdivision of the Parisian group, extending continuously throughout a great table-land intervening between the basin of the Seine and that of the Loire. The other example occurs in Italy, where strata containing many fossils similar to those of Bordeaux were observed by Bonelli and others in the environs of Turin, subjacent to strata belonging to the Subapennine group of Brocchi.
Value of Testacean Fossils in Classification.—It will be observed that in the foregoing allusions to organic remains, the testacea or the shell-bearing mollusca are selected as the most useful and convenient class for the purposes of general classification. In the first place, they are more universally distributed through strata of every age than any other organic bodies. Those families of fossils which are of rare and casual occurrence are absolutely of no avail in establishing a chronological arrangement. If we have plants alone in one group of strata and the bones of mammalia in another, we can draw no conclusion respecting the affinity or discordance of the organic beings of the two epochs compared; and the same may be said if we have plants and vertebrated animals in one series and only shells in another. Although corals are more abundant, in a fossil state, than plants, reptiles, or fish, they are still rare when contrasted with shells, because they are more dependent for their well-being on the constant clearness of the water, and are, therefore, less likely to be included in rocks which endure in consequence of their thickness and the copiousness of sediment which prevailed when they originated. The utility of the testacea is, moreover, enhanced by the circumstance that some forms are proper to the sea, others to the land, and others to fresh water. Rivers scarcely ever fail to carry down into their deltas some land-shells, together with species which are at once fluviatile and lacustrine. By this means we learn what terrestrial, fresh-water, and marine species coexisted at particular eras of the past: and having thus identified strata formed in seas with others which originated contemporaneously in inland lakes, we are then enabled to advance a step farther, and show that certain quadrupeds or aquatic plants, found fossil in lacustrine formations, inhabited the globe at the same period when certain fish, reptiles, and zoophytes lived in the ocean.
Among other characters of the molluscous animals, which render them extremely valuable in settling chronological questions in geology, may be mentioned, first, the wide geographical range of many species; and, secondly, what is probably a consequence of the former, the great duration of species in this class, for they appear to have surpassed in longevity the greater number of the mammalia and fish. Had each species inhabited a very limited space, it could never, when imbedded in strata, have enabled the geologist to identify deposits at distant points; or had they each lasted but for a brief period, they could have thrown no light on the connection of rocks placed far from each other in the chronological, or, as it is often termed, vertical series.
Classification of Tertiary Strata.—Many authors have divided the European Tertiary strata into three groups—lower, middle, and upper; the lower comprising the oldest formations of Paris and London before mentioned; the middle those of Bordeaux and Touraine; and the upper all those newer than the middle group.
In the first edition of the Principles of Geology, I divided the whole of the Tertiary formations into four groups, characterised by the percentage of recent shells which they contained. The lower tertiary strata of London and Paris were thought by M. Deshayes to contain only 3½ per cent of recent species, and were termed Eocene. The middle tertiary of the Loire and Gironde had, according to the specific determinations of the same conchologist, 17 per cent, and formed the Miocene division. The Subapennine beds contained 35 to 50 per cent, and were termed Older Pliocene, while still more recent beds in Sicily, which had from 90 to 95 per cent of species identical with those now living, were called Newer Pliocene. The first of the above terms, Eocene, is derived from eos, dawn, and cainos, recent, because the fossil shells of this period contain an extremely small proportion of living species, which may be looked upon as indicating the dawn of the existing state of the testaceous fauna, no recent species having been detected in the older or secondary rocks.
The term Miocene (from meion, less, and cainos, recent) is intended to express a minor proportion of recent species (of testacea), the term Pliocene (from pleion, more, and cainos, recent) a comparative plurality of the same. It may assist the memory of students to remind them, that the Miocene contain a minor proportion, and Pliocene a comparative plurality of recent species; and that the greater number of recent species always implies the more modern origin of the strata.
It has sometimes been objected to this nomenclature that certain species of infusoria found in the chalk are still existing, and, on the other hand, the Miocene and Older Pliocene deposits often contain the remains of mammalia, reptiles, and fish, exclusively of extinct species. But the reader must bear in mind that the terms Eocene, Miocene, and Pliocene were originally invented with reference purely to conchological data, and in that sense have always been and are still used by me.
Since the year 1830 the number of known shells, both recent and fossil, has largely increased, and their identification has been more accurately determined. Hence some modifications have been required in the classifications founded on less perfect materials. The Eocene, Miocene, and Pliocene periods have been made to comprehend certain sets of strata of which the fossils do not always conform strictly in the proportion of recent to extinct species with the definitions first given by me, or which are implied in the etymology of those terms.