It seems that, when the ancient lake of the Limagne first began to be filled with sediment, no volcanic action had yet produced lava and scoriæ on any part of the surface of Auvergne. No pebbles, therefore, of lava were transported into the lake—no fragments of volcanic rocks imbedded in the conglomerate. But at a later period, when a considerable thickness of sandstone and marl had accumulated, eruptions broke out, and lava and tuff were deposited, at some spots, alternately with the lacustrine strata. It is not improbable that cold and thermal springs, holding different mineral ingredients in solution, became more numerous during the successive convulsions attending this development of volcanic agency, and thus deposits of carbonate and sulphate of lime, silex, and other minerals were produced. Hence these minerals predominate in the uppermost strata. The subterranean movements may then have continued until they altered the relative levels of the country, and caused the waters of the lakes to be drained off, and the further accumulation of regular fresh-water strata to cease.
Lower Miocene Mammalia of the Limagne.—It is scarcely possible to determine the age of the oldest part of the fresh-water series of the Limagne, large masses both of the sandy and marly strata being devoid of fossils. Some of the lowest beds may be of Upper Eocene date, although, according to M. Pomel, only one bone of a Palæotherium has been discovered in Auvergne. But in Vélay, in strata containing some species of fossil mammalia common to the Limagne, no less than four species of Palæothere have been found by M. Aymard, and one of these is generally supposed to be identical with Palæotherium magnum, an undoubted Upper Eocene fossil, of the Paris gypsum, the other three being peculiar.
Not a few of the other mammalia of the Limagne belong undoubtedly to genera and species elsewhere proper to the Lower Miocene. Thus, for example, the Cainotherium of Bravard, a genus not far removed from the Anoplotherium, is represented by several species, one of which, as I learn from Mr. Waterhouse, agrees with Microtherium Renggeri of the Mayence basin. In like manner, the Amphitragulus elegans of Pomel, an Auvergne fossil, is identified by Waterhouse with Dorcatherium nanum of Kaup, a Rhenish species from Weissenau, near Mayence. A small species, also, of rodent, of the genus Titanomys of H. von Meyer, is common to the Lower Miocene of Mayence and the Limagne d’Auvergne, and there are many other points of agreement which the discordance of nomenclature tends to conceal. A remarkable carnivorous genus, the Hyænodon of Laizer, is represented by more than one species. The same genus has also been found in the Upper Eocene marls of Hordwell Cliff, Hampshire, just below the level of the Bembridge Limestone, and therefore a formation older than the Gypsum of Paris. Several species of opossum (Didelphis) are met with in the same strata of the Limagne. The total number of mammalia enumerated by M. Pomel as appertaining to the Lower Miocene fauna of the Limagne and Velay falls little short of a hundred, and with them are associated some large crocodiles and tortoises, and some Ophidian and Batrachian reptiles.
Lower Molasse of Switzerland.—The two upper divisions of the Swiss Molasse—the one fresh-water, the other marine—have already been described in the preceding chapter. I shall now proceed to treat of the third division, which is of Lower Miocene age. Nearly the whole of this Lower Molasse is fresh-water, yet some of the inferior beds contain a mixture of marine and fluviatile shells, the Cerithium margaritaceum, a well-known Lower Miocene fossil, being one of the marine species. Notwithstanding, therefore, that some of these Lower Miocene strata consist of old shingle-beds several thousand feet in thickness, as in the Rigi, near Lucerne, and in the Speer, near Wesen, mountains 5000 and 7000 feet above the sea, the deposition of the whole series must have begun at or below the sea-level.
The conglomerates, as might be expected, are often very unequal in thickness, in closely adjoining districts, since in a littoral formation accumulations of pebbles would swell out in certain places where rivers entered the sea, and would thin out to comparatively small dimensions where no streams or only small ones came down to the coast. For ages, in spite of a gradual depression of the land and adjacent sea-bottom, the rivers continued to cover the sinking area with their deltas; until finally, the subsidence being in excess, the sea of the Middle Molasse gained upon the land, and marine beds were thrown down over the dense mass of fresh-water and brackish-water deposit, called the Lower Molasse, which had previously accumulated.
Flora of the Lower Molasse.—In part of the Swiss Molasse, which belongs exclusively to the Lower Miocene period, the number of plants has been estimated at more than 500 species, somewhat exceeding those which were before enumerated as occurring in the two upper divisions. The Swiss Lower Miocene may best be studied on the northern borders of the Lake of Geneva, between Lausanne and Vevay, where the contiguous villages of Monod and Rivaz are situated. The strata there, which I have myself examined, consist of alternations of conglomerate, sandstone, and finely laminated marls with fossil plants. A small stream falls in a succession of cascades over the harder beds of pudding-stone, which resist, while the sandstone and plant-bearing shales and marls give way. From the latter no less than 193 species of plants have been obtained by the exertions of MM. Heer and Gaudin, and they are considered to afford a true type of the vegetation of the Lower Miocene formations of Switzerland—a vegetation departing farther in its character from that now flourishing in Europe than any of the higher members of the series before alluded to, and yet displaying so much affinity to the flora of Œningen as to make it natural for the botanist to refer the whole to one and the same Miocene period. There are, indeed, no less than 81 species of these Older Miocene plants which pass up into the flora of Œningen.
This fact is important as bearing on the propriety of classing the Lower Molasse of Switzerland as belonging to the Miocene rather than to the latter part of the Eocene period. There are, indeed, so many types among the fossils, both specific and generic, which have a wide range through the whole of the Molasse, that a unity of character is thereby stamped on the whole flora, in spite of the contrast between the plants of the uppermost and lowest formations, or between Oeningen and Monod. The proofs of a warmer climate, and the excess of arborescent over herbaceous plants, and of evergreen trees over deciduous species, are characters common to the whole flora, but which are intensified as we descend to the inferior deposits.
Nearly all the plants at Monod are contained in three layers of marl separated by two of soft sandstone. The thickness of the marls is ten feet, and vegetable matter predominates so much in some layers as to form an imperfect lignite. One bed is filled with large leaves of a species of fig (Ficus populina), and of a hornbeam (Carpinus grandis), the strength of the wind having probably been great when they were blown into the lake; whereas another contiguous layer contains almost exclusively smaller leaves, indicating, apparently, a diminished strength in the wind. Some of the upper beds at Monod abound in leaves of Proteaceæ, Cyperaceæ, and ferns, while in some of the lower ones Sequoia, Cinnamomum, and Sparganium are common. In one bed of sandstone the trunk of a large palm-tree was found unaccompanied by other fossils, and near Vevay, in the same series of Lower Miocene strata, the leaves of a palm of the genus Sabal (Fig. 151), a genus now proper to America, were obtained.
Among other genera of the same class is a Flabellaria occurring near Lausanne, and a magnificent Phœnicites allied to the date palm. When these plants flourished the climate must have been much hotter than now. The Alps were no doubt much lower, and the palms now found fossil in strata elevated 2000 feet above the sea grew nearly at the sea-level, as is demonstrated by the brackish-water character of some of the beds into which they were carried by winds or rivers from the adjoining coast.