The tribe of land quadrupeds most abundant in this formation is such as now inhabits alluvial plains and marshes, and the banks of rivers and lakes, a class most exposed to suffer by river inundations. Among these were several species of Palæotherium, a genus before alluded to. These were associated with the Anoplotherium, a tribe intermediate between pachyderms and ruminants. One of the three divisions of this family was called by Cuvier Xiphodon. Their forms were slender and elegant, and one, named Xiphodon gracile (Fig. 218), was about the size of the chamois; and Cuvier inferred from the skeleton that it was as light, graceful, and agile as the gazelle.

Fossil Footprints.—There are three superimposed masses of gypsum in the neighbourhood of Paris, separated by intervening deposits of laminated marl. In the uppermost of the three, in the valley of Montmorency, M. Desnoyers discovered in 1859 many footprints of animals occurring at no less than six different levels.[[8]] The gypsum to which they belong varies from thirty to fifty feet in thickness, and is that which has yielded to the naturalist the largest number of bones and skeletons of mammalia, birds, and reptiles. I visited the quarries, soon after the discovery was made known, with M. Desnoyers, who also showed me large slabs in the Museum at Paris, where, on the upper planes of stratification, the indented foot-marks were seen, while corresponding casts in relief appeared on the lower surfaces of the strata of gypsum which were immediately superimposed. A thin film of marl, which before it was dried and condensed by pressure must have represented a much thicker layer of soft mud, intervened between the beds of solid gypsum. On this mud the animals had trodden, and made impressions which had penetrated to the gypseous mass below, then evidently unconsolidated. Tracks of the Anoplotherium with its bisulcate hoof, and the trilobed footprints of Palæotherium, were seen of different sizes, corresponding to those of several species of these genera which Cuvier had reconstructed, while in the same beds were foot-marks of carnivorous mammalia. The tracks also of fluviatile, lacustrine, and terrestrial tortoises (Emys, Trionyx, etc.) were discovered, also those of crocodiles, iguanas, geckos, and great batrachians, and the footprints of a huge bird, apparently a wader, of the size of the gastornis, to be mentioned in the sequel. There were likewise the impressions of the feet of other creatures, some of them clearly distinguishable from any of the fifty extinct types of mammalia of which the bones have been found in the Paris gypsum. The whole assemblage, says Desnoyers, indicate the shores of a lake, or several small lakes communicating with each other, on the borders of which many species of pachyderms wandered, and beasts of prey which occasionally devoured them. The tooth-marks of these last had been detected by palæontologists long before on the bones and skulls of Paleotheres entombed in the gypsum.

Imperfection of the Record.—These foot-marks have revealed to us new and unexpected proofs that the air-breathing fauna of the Upper Eocene period in Europe far surpassed in the number and variety of its species the largest estimate which had previously been formed of it. We may now feel sure that the mammalia, reptiles, and birds which have left portions of their skeletons as memorials of their existence in the solid gypsum constituted but a part of the then living creation. Similar inferences may be drawn from the study of the whole succession of geological records. In each district the monuments of periods embracing thousands, and probably in some instances hundreds of thousands of years, are totally wanting. Even in the volumes which are extant the greater number of the pages are missing in any given region, and where they are found they contain but few and casual entries of the physical events or living beings of the times to which they relate. It may also be remarked that the subordinate formations met with in two neighbouring countries, such as France and England (the minor Tertiary groups above enumerated), commonly classed as equivalents and referred to corresponding periods, may nevertheless have been by no means strictly coincident in date. Though called contemporaneous, it is probable that they were often separated by intervals of many thousands of years. We may compare them to double stars, which appear single to the naked eye because seen from a vast distance in space, and which really belong to one and the same stellar system, though occupying places in space extremely remote if estimated by our ordinary standard of terrestrial measurements.

Calcaire silicieux, or Travertin inférieur, A.2 and 3, [Table]—This compact siliceous limestone extends over a wide area. It resembles a precipitate from the waters of mineral springs, and is often traversed by small empty sinuous cavities. It is, for the most part, devoid of organic remains, but in some places contains fresh-water and land species, and never any marine fossils. The calcaire siliceux and the calcaire grossier usually occupy distinct parts of the Paris basin, the one attaining its fullest development in those places where the other is of slight thickness. They are described by some writers as alternating with each other towards the centre of the basin, as at Sergy and Osny.

The gypsum, with its associated marls before described, is in greatest force towards the centre of the basin, where the calcaire grossier and calcaire silicieux are less fully developed.

Grès de Beauchamp, or Sables Moyens, A.4, [Table]—In some parts of the Paris basin, sands and marls, called the Grès de Beauchamp, or Sables moyens, divide the gypseous beds from the calcaire grossier proper. These sands, in which a small nummulite (N. variolaria) is very abundant, contain more than 300 species of marine shells, many of them peculiar, but others common to the next division.

MIDDLE EOCENE FORMATIONS OF FRANCE.

Calcaire Grossier, upper and middle, B.1, [Table]—The upper division of this group consists in great part of beds of compact, fragile limestone, with some intercalated green marls. The shells in some parts are a mixture of Cerithium, Cyclostoma, and Corbula; in others Limnea, Cerithium, Paludina, etc. In the latter, the bones of reptiles and mammalia, Palæotherium and Lophiodon, have been found. The middle division, or calcaire grossier proper, consists of a coarse limestone, often passing into sand. It contains the greater number of the fossil shells which characterise the Paris basin. No less than 400 distinct species have been procured from a single spot near Grignon, where they are imbedded in a calcareous sand, chiefly formed of comminuted shells, in which, nevertheless, individuals in a perfect state of preservation, both of marine, terrestrial, and fresh-water species, are mingled together. Some of the marine shells may have lived on the spot; but the Cyclostoma and Limnea, being land and fresh-water shells, must have been brought thither by rivers and currents, and the quantity of triturated shells implies considerable movement in the waters.

Nothing is more striking in this assemblage of fossil testacea than the great proportion of species referable to the genus Cerithium (see [p. 245]). There occur no less than 137 species of this genus in the Paris basin, and almost all of them in the calcaire grossier. Most of the living Cerithia inhabit the sea near the mouths of rivers, where the waters are brackish; so that their abundance in the marine strata now under consideration is in harmony with the hypothesis that the Paris basin formed a gulf into which several rivers flowed.