CANON TRISTRAM’S SUPPORT.
Canon Tristram’s paper, “On the Ornithology of Northern Africa” (Part iii., The Sahara, continued), was published in The Ibis, vol. i., October, 1859. The important conclusions alluded to above are contained at the end of the section upon the species of desert larks (pp. 429–433):
“Writing with a series of about 100 larks of various species from the Sahara before me, I cannot help feeling convinced of the truth of the views set forth by Messrs. Darwin and Wallace in their communications to the Linnean Society, to which my friend Mr. A. Newton last year directed my attention.... It is hardly possible, I should think, to illustrate this theory better than by the larks and chats of North Africa. In all these birds we trace gradual modifications of coloration and of anatomical structure, deflecting by very gentle gradations from the ordinary type; but when we take the extremes, presenting most marked differences.”
These differences, he concludes—
“have a very direct bearing on the ease or difficulty with which the animal contrives to maintain its existence.”
He then points out, upon the uniform surface of the desert it is absolutely necessary that animals shall be protected by their colour:
“Hence, without exception, the upper plumage of every bird, whether Lark, Chat, Sylvian, or Sandgrouse, and also the fur of all the small mammals, and the skin of all the Snakes and Lizards, is of one uniform isabelline or sand colour. It is very possible that some further purpose may be served by the prevailing colours, but this appears of itself a sufficient explanation. There are individual varieties in depth of hue among all creatures. In the struggle for life which we know to be going on among all species, a very slight change for the better, such as improved means of escaping from its natural enemies (which would be the effect of an alteration from a conspicuous colour to one resembling the hue of the surrounding objects), would give the variety that possessed it a decided advantage over the typical or other forms of the species. Now in all creatures, from Man downwards, we find a tendency to transmit individual varieties or peculiarities to the descendants. A peculiarity either of colour or form soon becomes hereditary when there are no counteracting causes, either from change of climate or admixture of other blood. Suppose this transmitted peculiarity to continue for some generations, especially when manifest advantages arise from its possession, and the variety becomes not only a race, with its variations still more strongly imprinted upon it, but it becomes the typical form of that country.”
Canon Tristram then points out the manner in which he imagines that one of the crested larks of the desert has been produced by the survival of the lightest coloured individuals, Galerida abyssinica only differing in this respect from G. cristata of Europe. Short-billed species of the same genus inhabiting hard rocky districts, and long-billed inhabiting loose sandy tracts have, he believes, been produced by the survival in each case of the forms of bill most suited to procure food:
“Here are only two causes enumerated which might serve to create as it were a new species from an old one, yet they are perfectly natural causes, and such as, I think, must have occurred, and are possibly occurring still. We know so very little of the causes which in the majority of cases make species rare or common, that there may be hundreds of others at work, some even more powerful than these, which go to perpetuate and eliminate certain forms ‘according to natural means of selection.’ But even these superficial causes appear sufficient to explain the marked features of the Desert races, which frequently approach so very closely the typical form, and yet possess such invariably distinctive characteristics, that naturalists seem agreed to elevate them to the rank of species.”
Although the author also declares his belief in the special creation of many species—a view put forward as possible by Darwin in the “Origin”[G]—and also believed in some direct influence of locality, climate, etc., the above quoted passages are a most complete acceptance of natural selection, at the same time affording excellent examples of its operation.