A primrose by a river's brim
A yellow primrose was to him,
And it was nothing more.
We all, I think, shared the same feeling, and found that the explanation of the flower then given, and to which I shall refer again, invested it with fresh interest and even with new beauty.
A regular flower, such, for instance, as a Geranium or a Pink, consists of four or more whorls of leaves, more or less modified: the lowest whorl is the Calyx, and the separate leaves of which it is composed, which however are sometimes united into a tube, are called sepals; (2) a second whorl, the corolla, consisting of coloured leaves called petals, which, however, like those of the Calyx, are often united into a tube; (3) of one or more stamens, consisting of a stalk or filament, and a head or anther, in which the pollen is produced; and (4) a pistil, which is situated in the centre of the flower, and at the base of which is the Ovary, containing one or more seeds.
Almost all large flowers are brightly coloured, many produce honey, and many are sweet-scented.
What, then, is the use and purpose of this complex organisation?
It is, I think, well established that the main object of the colour, scent, and honey of flowers is to attract insects, which are of use to the plant in carrying the pollen from flower to flower.
In many species the pollen is, and no doubt it originally was in all, carried by the air. In these cases the chance against any given grain of pollen reaching the pistil of another flower of the same species is of course very great, and the quantity of pollen required is therefore immense.
In species where the pollen is wind-borne as in most of our trees—firs, oaks, beech, ash, elm, etc., and many herbaceous plants, the flowers are as a rule small and inconspicuous, greenish, and without either scent or honey. Moreover, they generally flower early, so that the pollen may not be intercepted by the leaves, but may have a better chance of reaching another flower. And they produce an immense quantity of pollen, as otherwise there would be little chance that any would reach the female flower. Every one must have noticed the clouds of pollen produced by the Scotch Fir. When, on the contrary, the pollen is carried by insects, the quantity necessary is greatly reduced. Still it has been calculated that a Peony flower produces between 3,000,000 and 4,000,000 pollen grains; in the Dandelion, which is more specialised, the number is reduced to about 250,000; while in such a flower as the Dead-nettle it is still smaller.
The honey attracts the insects; while the scent and colour help them to find the flowers, the scent being especially useful at night, which is perhaps the reason why evening flowers are so sweet.
It is to insects, then, that flowers owe their beauty, scent, and sweetness. Just as gardeners, by continual selection, have added so much to the beauty of our gardens, so to the unconscious action of insects is due the beauty, scent, and sweetness of the flowers of our woods and fields.