Fig. 2.—Triarthrus Becki, Green.
A Trilobite of primitive type, showing its limbs and antennæ. (After Beecher.)
Figure 2, borrowed from Beecher,[4] shows the limbs of a species, not of the Lower Cambrian, but of a somewhat later formation. There can be no doubt, however, that those of earlier species were equally perfect, more especially as Triarthrus is an animal of an old type approaching to extinction in the age succeeding the Cambrian, and its representatives in the earlier and palmy days of the family could not have been inferior in organization. These creatures swarmed in every sea in the Cambrian period, and were represented by a great number of species, some of them of large size, others very small; some many-jointed, others few-jointed, and with a great variety of tubercles, spines, and other ornamental and protective parts. If we ask for their affinities and place in the great group of Crustacea, the answer must be that, while in some points allied to the higher forms, they approach most nearly to those which occupy a medium position in the class, and are, in fact, a composite type, presenting points of structure now distributed among different groups. If we ask for affinities with lower groups, we have to reply that their nearest allies in this direction are the bristle-footed marine worms; but there is a vast gap, both in the Cambrian and Modern seas, between any of these worms and the Crustacea, which, either as embryos or as adults, have any resemblance to them.
[4] American Journal of Science, 1896.
The Trilobites, after appearing in a great variety of generic and specific forms, and playing a most important part in their time, were not destined to continue beyond the Carboniferous period, and before that time they were beginning to give place to the Limuli, King-crabs, or Horseshoe-crabs, a few species of which continue on our coasts until the present time. In this limited duration the Trilobites present a strange contrast to certain shrimp-like Crustaceans, their contemporaries (the Phyllopods), which very closely resemble some still extant, and the same remark applies to swarms of little bivalve Crustaceans (Ostracods), which are still represented by hosts of modern species both in the sea and in the fresh waters. There is, however, a remarkable group of shrimp-like Crustaceans, represented in the modern world by only a few small species, which in the Cambrian age attained greater size, and constitute a very generalized type combining characters now found in lower and higher groups of Crustacea.
Hymenocaris vermicauda of Salter ([Fig. 3]) may serve to illustrate one of these primitive forms.
Fig. 3.—Hymenocaris vermicauda, Salter.
A Lower Cambrian Shrimp of generalized type. (After Salter.)
In point of fact, as Dr. Henry Woodward has shown in an able presidential address delivered to the Geological Society in 1895, at the base of the Lower Cambrian we still have several distinct groups of Crustacea; and if with some we were to hold them as traceable to one original form or to a worm-like ancestor, we must seek for this far back in those pre-Cambrian rocks in which we find no Crustaceans whatever. There is, it is true, no good reason to demand this; for whatever the cause, secondary or final, which produced any form of Crustacean in the Lower Cambrian, it might just as well have produced several distinct forms. Evolutionists seem to be somewhat unreasonable in demands of this kind, for any cause capable of originating a new form of living being, might have been operative at the same time in different localities and under somewhat diverse conditions, and may also have acted at different times. All imaginary lines of descent of animals are more or less subject to this contingency; and this may partly account for the great diversity in the lines of affiliation presented to us by evolutionists, which may in part have a basis in fact in so far as distinct varietal and racial forms are concerned, but may just as likely be entirely fallacious in the case of true species. In any case, in the lowest rocks into which we can trace Crustacea, we have already probably five of the orders into which their successors in the modern seas are divided by zoologists; and this is certainly a singular and suggestive fact, the significance of which we shall be better prepared to understand at a later stage of our investigation.
Allied in some respects to the Crustacea, though much lower in grade, are the marine Worms—a great and varied host—usually inhabiting the shallower parts of the ocean; though the 330 species collected by the Challenger expedition show that they also abound in those greater depths to which voyagers have only recently had access. Sea-worms seem thus to be able to live in all depths, as well as in all climates; and in accordance with this they abound in the oldest rocks, which are often riddled with the holes caused by their burrowing, or abundantly marked on the surfaces of the beds with their trails.