Fig. 8.—Diagrammatic section of two Laminæ of Cryptozoon, showing the Canals of the intermediate space, or Stroma (magnified).
Specimen in Peter Redpath Museum.
Of the whole we might perhaps have been able to secure at least 200 species even in one locality. The likelihood is that if there had been a collecting expedition like that of the Challenger in Early Cambrian times, it could have secured thousands of specific forms representing all the above types, more especially as we probably know very little of the softer and shell-less animals of these old seas, and there is some reason to believe that these may have been in greater proportion than in the present ocean.
In illustration of the richness of some parts of the lowest Cambrian sea, I may refer here to the large and beautifully illustrated Memoir of Walcott on the Lower Cambrian, containing fifty folio plates of species collected in a few districts of North America; and, as a minor example, to the contents of a loose boulder of limestone of that age, found at Little Metis on the Lower St. Lawrence, under the following circumstances ([Fig. 9]):—
Fig. 9.—Lower Cambrian Fossils found in a few cubic inches of limestone in a conglomerate at Little Metis; viz., Trilobites of genera Olenellus, Ptychoparia, Solenopleura, Protypus; Brachiopod of genus Iphidea; Pteropod of genus Hyolithes; Gastropod, genus Stenotheca; Sponge, undetermined.
Along what is now the valley of the Lower St. Lawrence and the gulf of the same name, there seem to have been deposited in the oldest Cambrian or Olenellus period beds of limestone rich in shells of marine animals and fragments of these. These can be seen in place in some parts of Newfoundland, and here and there on the hills bounding the St Lawrence River; but for the most part they have been swept away by the sea when these districts were being elevated to form parts of the American land. Their ruins appear as boulders and pebbles in thick beds of conglomerate or pudding-stone, constituting portions of the Upper Cambrian and Lower Ordovician series, which now occupy the south coast of the Lower St. Lawrence. In one of these boulders, less than a foot in diameter, removed from its hard matrix and carefully broken up, I found fragments representing eleven different species, of which no less than eight were trilobites, one a gastropod, one a brachiopod, and one probably a sponge—and this forms an interesting illustration of the number of species sometimes to be found in a limited space, and also of the great prevalence of the Trilobites in these beds. The statistics of these groups for North America, as given by Walcott, show 165 species belonging to all the groups enumerated above, and of these the Trilobita constitute one-third of the whole; so that the Olenellus Zone, as it has been called from one genus of these Crustaceans, might well be named the reign of Trilobites, unless, indeed, as the indications already referred to seem to show, giant cuttle-fishes, destitute of shells, were then the tyrants of the sea, but are represented only by the markings of their long and muscular arms on the soft sea mud while dashing after their Crustacean prey. What I desire, however, chiefly to emphasize is, that in the lowest beds of the Cambrian we have evidence of sea-bottoms swarming with representatives of all the leading types of marine invertebrate life, and therefore seem to be still far from the beginning of living things, if that was a slow and gradual process, rather than a sudden or rapid series of events.
PRE-CAMBRIAN LIFE
III
PRE-CAMBRIAN LIFE
H