The modern or human period of geology, that in which man and his contemporaries are certainly known to have inhabited the earth, was immediately preceded by an age of climatal refrigeration known as the Glacial or Ice age. This was further characterized not only by a prevalence of cold, unexampled so far as known either before or since, but by immense changes of the relative levels of sea and land, amounting, in some cases, at least, to several thousands of feet. The occurrence of these changes is clearly proved by the undoubted traces of the action of ice, whether land ice or floating ice, on all parts of our continents, half way to the equator, and by the occurrence of sea terraces and modern marine shells at high levels on mountains and table-lands. Perhaps we scarcely realize as we should the stupendous character of the changes involved in the driftage of heavy ice over our continents as far south as the latitude of 40°, in the deposit of boulders on hills several thousands of feet in height, and in the occurrence of shells of species still living in the sea, in beds raised to more than twelve hundred feet above its present level. Yet such changes must have occurred in the latest geological period immediately preceding that in which we live. Proceeding farther back in geological time, we find the still more extraordinary fact that in the middle and earlier Tertiary the northern hemisphere enjoyed a climate so much more mild than that which now prevails, that plants at present confined to temperate latitudes could flourish in Greenland and Spitzbergen.[183] The age in which we live is thus one of mediocrity, attaining neither to the Arctic rigour of the later Pleistocene, nor to the universal mildness of the preceding Miocene.
[183] As I have elsewhere shown, a warm climate in an Arctic region seems to have afforded the necessary conditions for the great colonizing floras of all geological periods.
The causes of these changes of climate we have discussed elsewhere. It remains for us now to consider the actual condition of our present continents, and the bearing of past conditions on the distribution of their living inhabitants.
In speaking of continents and islands, it may be as well to remark at the outset that all the land existing, or which probably has at any time existed, consists of islands great or small. It is all surrounded by the ocean. Two of the greater masses of land are, however, sufficiently extensive to be regarded as continents, and from their very extent and consequent permanence may be considered as the more special homes of the living beings of the land. Two other portions of land, Australia and the Antarctic polar continent, may be regarded either as smaller continents or large islands, but partake of insular rather than continental characters in their animals and plants. All the other portions of land are properly islands; but while these islands, and more especially those in mid-ocean, cannot be regarded as the original homes of many forms of life, we shall find that they have a special interest as the shelters and refuges of numerous very ancient and now decaying species.
The two great continents of America and Eurasia have been the most permanent portions of the land throughout geological time, some parts of them having always been above water, probably from the Laurentian age downward, though at various times they have been reduced to little more than groups of islands. On them, and more especially in their more northern parts, in which the long continuance of daylight in summer seems in warm periods to have been peculiarly favourable to the introduction of new vegetable and animal forms, and to the giving to them that vigour necessary for active colonization, have originated the greater number of the inhabitants of the land.
Regarded as portions of the earth's crust, the continents are areas in which the lateral thrust, caused by the secular contraction of the interior of the earth and unequal settlement of the crust, has ridged up and folded the rocks, producing mountain chains. This process began in the earliest geological periods, and has been repeated at long intervals, the original lines of folding guiding those formed in each new thrust proceeding from the broad oceanic areas. Along the ridges thus produced, and in the narrower spaces between them, the greater part of the sediment carried by water was laid down, thus producing plateaus in connection with the mountain-chains, while the weight of new sediments and the removal of matter from other areas by denudation, have been constantly producing local depression and elevation. The tendency of the ocean to be thrown toward the poles by the retardation of the earth's rotation, alternating with great collapses of the crust at the equator proceeding from the same cause, along with the secular cooling, have produced alternate submergence and emergence of these plateaus. This has been further complicated by the constant tendency of the Arctic and Antarctic currents, aided by ice, to drift solid materials, set free by the vast denuding action of frost, from the polar to the temperate regions, and by the further tendency of animal life to heap up calcareous accumulations under the warm waters of the tropical regions. All these changes, as already stated, have conspired to modify the directions of the great oceanic currents, and to produce vicissitudes of climate under which animals and plants have been subjected in geological time to those migrations, extinctions, and renovations of which their fossil remains and present distribution afford evidence.
Still, it is true that throughout the whole of these great mutations, since the beginning of geological history, there seems never to have been any time when the ocean so regained its dominion as to produce a total extinction of land life; still less was there any time when the necessary conditions of all the various forms of marine life failed to be found; nor was there any climatal change so extreme as to banish any of the leading forms of life from the earth. To geologists it is not necessary to say that the conclusions sketched above are those that have been reached as the results of long and laborious investigation, and which have been illustrated and established by Lyell, Dana, Wallace,[184] and many other writers.[185] Let us now place beside them some facts as to the present distribution of life, and of the agencies which influence it.
[184] Wallace, "Geographical Distribution of Animals" and "Island Life." Second edition.
[185] The writer has endeavoured to popularize these great results of geology in his work, the "Story of the Earth." Ninth Edition. London, 1887. They are often overlooked by specialists, and by compilers of geological manuals.
Just as political geography sometimes presents boundaries not in accordance with the physical structure of countries, so the distribution of animals and plants shows many peculiar and unexpected features. Hence naturalists have divided the continents into what Sclater has called zoological regions, which are, so to speak, the great empires of animal life, divisible often by less prominent boundaries into provinces. In vegetable life similar boundaries may be drawn, more or less coincident with the zoological divisions. Zoologically, North America and Greenland may be regarded as one great region, the Nearctic, or new Arctic, the prefix not indicating that the animals are newer than those of the old world, which is by no means the case. South America constitutes another region the Neotropical. If now we turn to the greater Eurasian continent, with its two prolongations to the south in Africa and Australia, we shall find the whole northern portion, from the Atlantic to the Pacific, constituting one vast region of animal life, the Palearctic, which also includes Iceland and a strip across North Africa. Africa itself, with Madagascar, whose allegiance is, however, only partial, constitutes the Ethiopian region. India, Burmah, the south of China, and certain Asiatic islands form the Oriental region. Australia, New Guinea, and the Polynesian islands constitute the Australian region. All of these regions may in a geological point of view be considered as portions of old and permanent continental masses, which, though with movements of elevation and depression, have continued to exist for vast periods. Some of them, however, seem to have enjoyed greater immunity from causes of change than others, and present, accordingly, animals and plants having, geologically speaking, an antique aspect in comparison. In this sense the Australian province may be regarded as the oldest of all in the facies of its animal forms, since creatures exist there of genera and families which have very long ago become extinct everywhere else. Next in age to this should rank the Neotropical or South American region, which, like Australia, presents many low and archaic forms of animal life. The Ethiopian region stands next to it in this, the Oriental and Nearctic next, and last and most modern in its aspect is the great Palearctic region, to which man himself belongs, and the animals and plants of which vindicate their claims to youth by that aggressive and colonizing character already referred to, and which has enabled them to spread themselves widely over the other regions, even independently of the influence of man. On the other hand, the animals and plants of the Australian and South American regions show no such colonizing tendency, and can scarcely maintain themselves against those of other regions when introduced among them. Thus we have at once in these continental regions a great and suggestive example of the connection of geographical and geological distribution, the details of which are of the deepest interest, and have not yet been fully worked out. One great principle is, however, sufficiently established; namely, that the northern regions have been the birthplace of new forms of land life, whence they have extended themselves to the south, while the comparative isolation and equable climate of the South American and Australian regions have enabled them to shelter and retain the old moribund tribes.