Here we may first remark that it is not so much the physical record of geology that is imperfect as the organic record. Ever since the time of Hutton and Playfair we have learned that the processes of mineral detrition and deposition are continuous, and have been so throughout geological time. The erosion of the land is constantly going on, every shower carries its tribute of earthy matter toward the sea, and every wave that strikes against a beach or cliff does some work toward the grinding of shells, pebbles or stone. Thus, everywhere around our continents there is a continuous deposition of beds of earthy matter, and it is this which, when elevated into new land, has given us our chronological series of geological formations. True, the elevating process is not continuous, but, so far as we know, intermittent; but it has been so often repeated that we have no reason to doubt that the wasting continents afford a complete series of aqueous deposits, since the time when the dry land first appeared.
In recent years the Challenger expedition and similar dredgings have informed us of still another continuity of deposition in the depths of the ocean. There, where no detritus from the land, or only a very little fine volcanic ash or pumice has ever reached, we have, going on from age to age, a deposit of the hard parts of abyssal animals and of those that swim in the open sea; so that if it were possible to bore or sink a shaft in some parts of the ocean, we should find not only a continuous bed, but a continuous series of pelagic life from the Laurentian to the present day. Thus we have continuous physical records, could we but reach or completely put them together, and eliminate the disturbing influence of merely local vicissitudes. It is when we begin to search the geological formations for fossils, that imperfection in our record first becomes painfully manifest.
In the case of many groups of marine animals, as, for example, the shell-fish and the corals, and I may add the bivalve crustaceans, so admirably worked up by my friend Prof. Rupert Jones, we have very complete series. With the and snails the case is altogether different. As stated in another paper of this series, a few species of these animals appear in the later Palæozoic age, and after that they have no successors known to us in all the great periods covered by the Permian, the Trias, and the earlier Jurassic. A few air-breathing water-snails appear in the upper Jurassic, and true land snails are not met with again until the Tertiary. Were there no land snails in this vast lapse of time? Have we two successive creations, so to speak, of these creatures at distant intervals? Were they only diminished in numbers and distribution in the intervening time? Is the hiatus owing merely to the unlikelihood of such shells being preserved? Or is it owing to the lack of diligence and care in collecting?
In this particular case we are, no doubt, disposed to say that the series must have been continuous. But we cannot be sure of this. In whatever way a few species of land snails were so early introduced in the time of the Devonian or of the Coal formation, if from physical vicissitudes or lack of proper pabulum they became extinct, there is no reason known to us why, when circumstances again became favourable, they should not be reintroduced in the same manner as at first, whether by development from allied types or otherwise. The fact that the few Devonian and Carboniferous species are very like those that still exist, perhaps makes against this supposition, but does not exclude it. If we suppose that new forms of life of low grade are introduced from time to time in the course of the geological ages, and if we adopt the Darwinian hypothesis of evolution, we arrive, as Naegeli has so well pointed out, at the strange paradox, that the highest forms of life must be the oldest of all, since they will be the descendants of the earliest of the lower animals, whereas the animals now of low grade may have been introduced later, and may not have had time to improve. But all our attempts to reduce nature to one philosophic expression necessarily lead to such paradoxes.
On the other hand, the chances of the preservation of land snails in aqueous deposits are vastly less than those in favour of the preservation of aquatic species. The first Carboniferous species found[15] had been preserved in the very exceptional circumstances afforded by the existence of hollow trunks of Sigillariæ on the borders of the Coal formation flats, and the others subsequently found were in beds no doubt receiving the drainage of neighbouring land areas. Still it is not uncommon on the modern sea-shore, anywhere near the mouths of rivers, to find a few fresh-water shells here and there. The carbonaceous beds of the Trias, the fossil soils of the Portland series, the estuarine Wealden beds would seem to be as favourably situated as those of the coal formation for preserving land shells, though possibly the flora of the Mesozoic was less suitable for feeding such creatures than that of the Coal period, and they may consequently have become few and local. After all, perhaps more diligent collecting and more numerous collectors might succeed, and may succeed in the future, in filling this and similar gaps.
[15] Pupa vetusta of the Nova Scotia coal formation.
It is a great mistake to suppose that discoveries of this kind are made by chance. It is only by the careful and painstaking examination of much material that the gaps in the geological record can be filled up, and I propose in the sequel of this article to note a few instances, in a country where the range of territory is altogether out of proportion to the number of observers, and which have come within my own knowledge.
It was not altogether by accident that Sir C. Lyell and the writer discovered a few reptilian bones and a land-snail in breaking up portions of the material filling an erect Sigillaria in the South Joggins coal measures. We were engaged in a deliberate survey of the section, to ascertain as far as might be the conditions of accumulation of coal, and one point which occurred to us was to inquire as to the circumstances of preservation of stumps of forest trees in an erect position, to trace their roots into the soils on which they stood, and to ascertain the circumstances in which they had been buried, had decayed, and had been filled with mineral matter. It was in questioning these erect trees on such subjects and this not without some digging and hammering that we made the discovery referred to.
But we found such remains only in one tree, and they were very imperfect, and indicated only two species of batrachians and one land-snail. There the discovery might have rested. But I undertook to follow it up. In successive visits to the coast, a large number of trees standing in the cliff and reefs, or fallen to the shore, were broken up and examined, the result being to discover that, with one unimportant exception, the productive trees were confined to one of the beds at Coal Mine Point, that from which the original specimens had been obtained. Attention was accordingly concentrated on this, and as many as thirty trees were at different times extracted from it, of which rather more than one-half proved more or less productive. By these means bones representing about sixty specimens and twelve species were extracted, besides numerous remains of land shells, millipedes, and scorpions. In this way a very complete idea was obtained of the land life, or at least of the smaller land animals, of this portion of the coal formation of Nova Scotia. It is not too much to say that if similar repositories could be found in the succeeding formations, and properly worked when found, our record of the history of land quadrupeds might be made very complete.
When in 1855 I changed my residence from Nova Scotia to Montreal, and so was removed to some distance from the carboniferous rocks which I had been accustomed to study, I naturally felt somewhat out of place in a Cambro-Silurian district, more especially as my friend Billings had already almost exhausted its fossils. I found, however, a congenial field in the Pleistocene shell beds; more especially as I had given some attention to recent marine animals when on the sea coast. The very perfect series of Pleistocene deposits in the St. Lawrence valley locally contain marine shells from the bottom of the till or boulder clay up to the overlying sands and gravels. The assemblage was a more boreal one than that on the coast of Nova Scotia, though many of the species were the same, and both the climatal and bathymetrial conditions differed in different parts of the Pleistocene beds themselves. The gap in the record here could at that time be filled up only by collecting recent shells. In addition to what could be obtained by exchanging with naturalists who had collected in Greenland, Labrador, and Norway, I employed myself, summer after summer, in dredging both on the south and north shore of the St. Lawrence, until able at length to discover in a living state, but under different conditions as to temperature and depth, nearly every species found in the beds on the land, from the lower boulder clay to the top of the formation, and from the sea-level to the beds six hundred feet high on the hills. Not only so: I could ascertain in certain places and conditions all the peculiar varieties of the species, and the special modes of life which they indicated. Thus, in the cases of the Peter Redpath Museum, and in notes on the Post-pliocene of Canada, the gap between the Modern and the Glacial age was completely filled up in so far as Canadian marine species are concerned. The net result was, as I have elsewhere stated, that no change other than varietal had occurred.