[88] American Journal of Science, xvi., 1878.
"I can only say, at large, that the same species (of Tertiary fossil plants) have been found all round the world; that the richest and most extensive finds are in Greenland; that they comprise most of the sorts which I have spoken of, as American trees which once lived in Europe—Magnolias, Sassafras, Hickories, Gum-trees, our identical Southern Cypress (for all we can see of difference), and especially Sequoias, not only the two which obviously answer to the two Big-trees now peculiar to California, but several others; that they equally comprise trees now peculiar to Japan and China—three kinds of Gingko-trees, for instance, one of them not evidently distinguishable from the Japan species which alone survives; that we have evidence, not merely of Pines and Maples, Poplars, Birches, Lindens, and whatever else characterize the temperate-zone forests of our era, but also of particular species of these, so like those of our own time and country, that we may fairly reckon them as the ancestors of several of ours. Long genealogies always deal more or less in conjecture; but we appear to be within the limits of scientific inference when we announce that our existing temperate trees came from the north, and within the bounds of high probability when we claim not a few of them as the originals of present species. Remains of the same plants have been found fossil in our temperate region, as well as in Europe."
Between 1860 and 1870 the writer was engaged in working out all that could be learned of the Devonian plants of Eastern America, the oldest known flora of any richness, and which consists almost exclusively of gigantic, and to us grotesque, representatives of the Club mosses, Ferns, and Mare's-tails, with some trees allied to the Cycads and Pines. In this pursuit nearly all the more important localities were visited, and access was had to the large collections of Professor Hall and Professor Newberry in New York and Ohio, as well as to those of the Geological Survey of Canada, and to those made in the remarkable plant-bearing beds of St. John, New Brunswick, by Messrs. Matthew and Hartt. In the progress of these researches, which developed an unexpectedly rich assemblage of species, the northern origin of this old flora seemed to be established by its earlier culmination in the north-east, in connection with the growth of the American land to the southward, which took place after the great Upper Silurian subsidence, by elevations which began in the north, while those portions of the continent to the south-west still remained under the sea.
When, in 1870, the labours of those ten years were brought before the Royal Society of London, in the Bakerian Lecture of that year, and in a memoir illustrating no less than one hundred and twenty-five species of plants older than the great Carboniferous system, these deductions were stated in connection with the conclusions of Hall, Logan, and Dana, as to the distributions of sediment along the north-east side of the American continent, and the anticipation was hazarded that the oldest Palæozoic floras would be discovered to the north of Newfoundland. Mention was also made of the apparent earlier and more copious birth of the Devonian flora in America than in Europe, a fact which is itself connected with the greater northward extension of this continent.
Unfortunately the memoir containing these results was not published by the Royal Society, and its publication was secured in a less perfect form only in the reports of the Geological Survey of Canada. The part of the memoir relating to Canadian fossil plants, with a portion of the theoretical deductions, was published in a report issued in 1871.[89] In this report the following language was used:—
[89] "Fossil Plants of the Devonian and Upper Silurian Formations of Canada," pp. 92, twenty plates. Montreal, 1871.
"In Eastern America, from the Carboniferous period onward, the centre of plant distribution has been the Appalachian chain. From this the plants and sediments extended westward in times of elevation, and to this they receded in times of depression. But this centre was non-existent before the Devonian period, and the centre of this must have been to the north-east, whence the great mass of older Appalachian sediment was derived. In the Carboniferous period there was also an eastward distribution from the Appalachians, and links of connection in the Atlantic bed between the floras of Europe and America. In the Devonian such connection can have been only far to the north-east. It is therefore in Newfoundland, Labrador, and Greenland that we are to look for the oldest American flora, and in like manner on the border of the old Scandinavian nucleus for that of Europe."
"Again, it must have been the wide extension of the sea of the Corniferous limestone that gave the last blow to the remaining flora of the Lower Devonian: and the re-elevation in the middle of that epoch brought in the Appalachian ridges as a new centre, and established a connection with Europe which introduced the Upper Devonian and Carboniferous floras. Lastly, from the comparative richness of the later Erian[90] flora in Eastern America, especially in the St. John beds, it might be a fair inference that the north-eastern end of the Appalachian ridge was the original birthplace or centre of creation of what we may call the later Palæozoic flora, or a large part of that flora."
[90] The term Erian is used as synonymous with Devonian, and probably should be preferred to it, as pointing to the best development of this formation known, which is on the shores of Lake Erie.
When my paper was written I had not seen the account published by the able Swiss palæobotanist Heer, of the remarkable Devonian flora of Bear Island, near Spitzbergen.[91] From want of acquaintance with the older floras of America and Western Europe, Heer fell into the unfortunate error of regarding the Bear Island plants as Lower Carboniferous, a mistake which his great authority has tended to perpetuate, and which has even led to the still graver error of some European geologists, who do not hesitate to regard as Carboniferous the fossil plants of the American deposits from the Hamilton to the Chemung groups inclusive, though these belong to formations underlying the oldest Carboniferous, and characterized by animal remains of unquestioned Devonian age. In 1872 I addressed a note to the Geological Society of London on the subject of the so-called "Ursa stage" of Heer, showing that though it contained some forms not known at so early a date in temperate Europe, it was clearly Devonian when tested by North American standards; but that in this high latitude, in which, for reasons stated in the report above referred to, I believed the Devonian plants to have originated, there might be an intermixture of the two floras. But such a mixed group should in that latitude be referred to a lower horizon than if found in temperate regions.