It has often happened to geologists, as to other explorers of new regions, that footprints on the sand have guided them to the inhabitants of unknown lands, and such footprints, proverbially perishable, may be so preserved by being filled up with matter deposited in them as to endure for ever. This we may see to-day in the tracks of sandpipers and marks of rain-drops preserved in the layers of alluvial mud deposited by the tides of the Bay of Fundy, and which, if baked or hardened by pressure, might become imperishable, like the inscriptions of the old Chaldeans on their tablets of baked clay. The first trace ever observed of reptiles in the Carboniferous system consisted of a series of small but well-marked footprints found by Sir W. E. Logan, in 1841, in the lower coal measures of Horton Bluff, in Nova Scotia; and as the authors of most of our general works on geology have hitherto, in so far as I am aware, failed to do justice to this discovery, I shall notice it here in detail. In the year above mentioned, Sir William, then Mr. Logan, examined the coal fields of Pennsylvania and Nova Scotia, with the view of studying their structure, and extending the application of the discoveries as to beds with roots, or Stigmaria underclays, which he had made in the Welsh coal fields. On his return to England he read a paper on these subjects before the Geological Society of London, in which he noticed the subject of reptilian footprints at Horton Bluff. The specimen was exhibited at the meeting of the Society, and was, I believe, admitted, on the high authority of Prof. Owen, to be probably reptilian. Unfortunately Sir William's paper appeared only in abstract in the Transactions; and in this abstract, though the footprints are mentioned, no opinion is expressed as to their nature. Sir William's own opinion is thus stated in a letter to me, dated June, 1843, when he was on his way to Canada, to commence the survey which has since developed so astonishing a mass of geological facts.

Footprints of Hylopus Logani, Dawson, Lower Carboniferous, Nova Scotia.
Natural size and reduced.

These footprints were the first indications of Carboniferous land vertebrates ever observed; they were probably made by a Microsaurian and one of the earliest species of this type. They show a remarkable length of stride and development of limb.

"Among the specimens which I carried from Horton Bluff, one is of very high interest. It exhibits the footprints of some reptilian animal. Owen has no doubt of the marks being genuine footprints. The rocks of Horton Bluff are below the gypsum of that neighbourhood; so that the specimen in question (if Lyell's views are correct[125]) comes from the very bottom of the coal series, or at any rate very low down in it, and demonstrates the existence of reptiles at an earlier epoch than has hitherto been determined; none having been previously found below the magnesian limestone, or, to give it Murchison's new name, the 'Permian era.'"

[125] Sir Charles Lyell had then just read a paper announcing his discovery that the gypsiferous system of Nova Scotia is Lower Carboniferous, in which he mentions the footprints referred to, as being reptilian.

This extract is of interest, not merely as an item of evidence in relation to the matter now in hand, but as a mark in the progress of geological investigation. For the reasons above stated, the important discovery thus made in 1841, and published in 1842, was overlooked; and the discovery of reptilian bones by Von Dechen, at Saarbruck, in 1844, and that of footprints by Dr. King in the same year, in Pennsylvania, have been uniformly referred to as the first observations of this kind. Insects and Arachnidans, it may be observed, had previously been discovered in the coal formation in Europe.

The original specimen of these footprints is still in the collection of the Geological Survey of Canada, and a cast which Logan kindly presented to me is exhibited in the Peter Redpath Museum of McGill University. It is a slab of dark-coloured sandstone, glazed with fine clay on the surface; and having a series of seven footprints in two rows, distant about three inches; the distance of the impressions in each row being three or four inches, and the individual impressions about one inch in length. They seem to have been made by the points of the toes, which must have been armed with strong and apparently blunt claws, and appear as if either the surface had been somewhat firm, or the body of the animal had been partly water-borne. In one place only is there a distinct mark of the whole foot, as if the animal had exerted an unusual pressure in turning or stopping suddenly. One pair of feet—the fore feet, I presume—appear to have had four toes touching the ground; the other pair show only three or four, and it is to be observed that the outer toe, as in the larger footprints discovered by Dr. King, projects in the manner of a thumb, as in the cheirotherian tracks of the Trias. At a later date another series of footprints, possibly of the same animal, was obtained at the same place by Prof. Elder, and is now in the Peter Redpath Museum. Each foot in this shows five toes, and it is remarkable that the animal was digitigrade and took a long step for its size, indicating a somewhat high grade of quadrupedal organization. No mark of the tail or belly appears. The impressions are such as may have been made by animals similar to some of those to be described in the sequel.

Shortly afterward, Dr. Harding, of Windsor, when examining a cargo of sandstone which had been landed at that place from Parrsboro', found on one of the slabs a very distinct series of footprints, each with four toes, and a trace of the fifth. Dr. Harding's specimen is now in the museum of King's College, Windsor. Its impressions are more distinct, but not very different otherwise from those above described, as found at Horton Bluff. The rocks at that place are probably of nearly the same age with those of Parrsboro'. I afterward examined the place from which this slab had been quarried, and satisfied myself that the beds are Carboniferous, and probably Lower Carboniferous. They were ripple-marked and sun-cracked, and I thought I could detect some footprints, though more obscure than those in Dr. Harding's slab. Similar footprints are also stated to have been found by Dr. Gesner, at Parrsboro'. All of these were from the lowest beds of the Carboniferous system.