Another link in the chain of being remains to be noticed here. In the Laurentian limestones we meet with numerous minute spherical bodies and groups of spheres with calcareous tubulated tests.[7] These may either be small Foraminiferæ, distinct from Eozoon, or may be germs or detached cells from its surface. Similar bodies are found in the lower part of the Siluro-Cambrian, in the Quebec group at Point Levis; and there they are filled with a species of glauconite constituting a sort of greensand rock. Still higher, in the Carboniferous, there are very numerous species of Foraminifera, presenting forms very similar to those in the modern seas, so that in the smaller shells of this group we seem to have evidence of a continuous series all the way from the Laurentian to the present time. The greater laminated forms co-exist with these up to the Eocene Tertiary. Throughout the whole of geological time—from the formation of the Laurentian limestones to that of the chalky ooze accumulating in the modern ocean—these humble creatures have been among the chief instruments in seizing on the calcareous matter of the waters and depositing it in the form of limestone.
Fig. 27.—Foraminiferal Rock Builders, in the Cretaceous and Eocene.
a, Nummulites lævigata—Eocene. b, The same, showing chambered interior. c, Milioline limestone, magnified—Eocene, Paris. d, Hard Chalk, section magnified—Cretaceous.
I have said nothing of the development of higher forms of animal life from Eozoon, simply because I know nothing of it. We shall see in the next chapter that these are introduced seemingly in an independent manner. We may be content to trace foraminiferal life along its own line of development, waxing and waning, but ever confined within the same general boundaries, from the Laurentian to the present time. It is likely that if, in any of the ages constituting this vast lapse of time, a dredge had been dropped into the depths of ocean, it would have brought up Foraminifera not essentially different in form and structure. If any one asks to what extent the successive species constituting this almost endless chain may be descendants one of the other, we have no absolutely certain information to give. On the one hand, it is not inconceivable that such forms as Stromatopora or Nummulina may have descended from Eozoon. On the other hand, it is equally conceivable that the same power which produced Eozoon at first, whether from dead matter or from some unknown lower form of life, may have repeated the process in later times with modifications. In any case it is probable that the Foraminifera have experienced alternations of expansion and shrinkage, of elevation and decadence, in the lapse of geological time. There were times in which many new forms swarmed into existence, and times in which old forms were becoming extinct without being replaced by others. In so far as the areas of the continents and the adjacent waters are concerned, those periods when the land was subsiding under the ocean must have been their times of prosperity, those in which the crust of the earth shrunk and raised up large areas of land must have been their times of decay. Still this lowest form of animal life has never perished, but has always found abundant place for itself, however pressed by physical change and by the introduction of higher beings.