Inhabitants of the English Seas in the Age of Reptiles.
Pliosaurus, Ichthyosaurus, Plesiosaurus, Mososaurus, and Teleosaurus.
CHAPTER VII.
the empire of the great reptiles.
ad we lived in the Carboniferous period, we might have supposed that the line of the great Labyrinthodont Batrachians would have been continued onward and elevated, perhaps, in the direction of the Mammalia, to which some features of their structure point. But we should have been mistaken in this. The Labyrinthodonts, it is true, extend into the Trias; but there is perhaps a sign of their coming degradation in the appearance in the Permian of the first known mud-eel, a humble Batrachian form allied to the Newts and Water-lizards.[62] Their special peculiarities are dropped in the Mesozoic in favour of those of certain small and feeble lizard-like animals, appearing first in the Carboniferous, and more manifestly in the Permian, and which are the true forerunners, though they can scarcely be the ancestors, of the magnificent reptilian species of the Mesozoic, which have caused this period to be called “the age of reptiles.”
The leading reptilian animal from the European Permian has long been the Proterosaurus, from the copper slates of Thuringia ([Fig. 141]), a reptile of lizard-like form, with well-developed limbs, and attaining a length of three or four feet. It resembles more nearly those large modern lizards known as “Monitors,” than any other existing form. The fore-limb represented in the figure foreshadows very closely the bones of the human arm and hand. Besides this we find in the Permian certain lizards (Theriodonts of Owen) which present the remarkable and advanced peculiarity already predicted by some Carboniferous Microsauria,[63] of having distinct canine teeth, producing a division into incisors, canines, and molars, in the manner of the Carnivorous quadrupeds, which they seem also to have resembled in some other parts of their skeletons. It is not impossible that the footprints in the Permian sandstones of Scotland, which have been referred to tortoises, were those of animals of this type. Cope has recently described from the Permian of Texas a number of reptiles which have the complex dentition of the Theriodonts, and others which simulate that of Herbivorous mammals, by the possession of flat grinding teeth supposed to be adapted to vegetable food.[64] The teeth of all these Permian reptiles were set in sockets, also an advanced peculiarity. Thus already in the Permian, before the final decadence of the Carboniferous flora, and while the Palæozoic invertebrates still lingered in the sea, the age of reptiles dawned, and gave promise of its future greatness by the assumption on the part of reptilian species of structures now limited to the Mammalia.