The Geological History of Plants

by Sir John William Dawson

When, in 1870, the labours of those ten years were brought before the Royal Society of London, in the Bakerian lecture of that year, and in a memoir illustrating no less than one hundred and twenty-five species of plants older than the great Carboniferous system, these deductions were stated in connection with the conclusions of Hall, Logan, and Dana, as to the distribution of sediment along the northeast side of the American continent, and the anticipation was hazarded that the oldest Palæozoic floras would be discovered to the north of Newfoundland. Mention was also made of the apparent earlier and more copious birth of the Devonian flora in America than in Europe, a fact which is itself connected with the greater northward extension of this continent.

The memoir containing these results was not published by the Royal Society, but its publication was secured in a less complete form in the reports of the “Geological Survey of Canada.” The part of the memoir relating to Canadian fossil plants, with a portion of the theoretical deductions, was published in a report issued in 1871.[EG] In this report the following language was used:

[EG] “Fossil Plants of the Devonian and Upper Silurian Formations of Canada,” pp. 92, twenty plates, Montreal, 1871.

"In eastern America, from the Carboniferous period onward, the centre of plant distribution has been the Appalachian chain. From this the plants and sediments extended westward in times of elevation, and to this they receded in times of depression. But this centre was nonexistent before the Devonian period, and the centre for this must have been to the northeast, whence the great mass of older Appalachian sediment was derived. In the Carboniferous period there was also an eastward distribution from the Appalachians, and links of connection in the Atlantic bed between the floras of Europe and America. In the Devonian such connection can have been only far to the northeast. It is therefore in Newfoundland, Labrador, and Greenland that we are to look for the oldest American flora, and in like manner on the border of the old Scandinavian nucleus for that of Europe.

"Again, it must have been the wide extension of the sea of the corniferous limestone that gave the last blow to the remaining flora of the Lower Devonian; and the re-elevation in the middle of that epoch brought in the Appalachian ridges as a new centre, and established a connection with Europe which introduced the Upper Devonian and Carboniferous floras. Lastly, from the comparative richness of the later Erian[EH] flora in eastern America, especially in the St. John beds, it might be a fair inference that the northeastern end of the Appalachian ridge was the original birthplace or centre of creation of what we may call the later Palæozoic flora, or of a large part of that flora."

[EH] See [pages 107 and 108].

When my paper was written I had not seen the account published by the able Swiss palæobotanist Heer, of the remarkable Devonian flora of Bear Island, near Spitzbergen.[EI] From want of acquaintance with the older floras of America and western Europe, Heer fell into the unfortunate error of regarding the whole of Bear Island plants as Lower Carboniferous, a mistake which his great authority has tended to perpetuate, and which has even led to the still graver error of some European geologists, who do not hesitate to regard as Carboniferous the fossil plants of the American deposits from the Hamilton to the Chemung groups inclusive, though these belong to formations underlying the oldest Carboniferous, and characterised by animal remains of unquestioned Devonian age. In 1872 I addressed a note to the Geological Society of London on the subject of the so-called “Ursa stage” of Heer, showing that, though it contained some forms not known at so early a date in temperate Europe, it was clearly, in part at least, Devonian when tested by North American standards; but that in this high latitude, in which, for reasons stated in the report above referred to, I believed the Devonian plants to have originated, there might be an intermixture of the two floras. But such a mixed group should in that latitude be referred to a lower horizon than if found in temperate regions. Dr. Nathorst, as already stated, has recently obtained new facts which go to show that plants of two distinct horizons may have been intermixed in the collections submitted to Heer.

[EI] “Transactions of the Swedish Academy” 1871; “Journal of the London Geological Society,” vol. xxviii.