[EW] It seems certain that the faunæ of the old limestones, like the Trenton, Niagara, Lower Helderberg, and Corniferous, belong to warm and sheltered sea areas, and that those rich in graptolites and trilobites, enclosed in muddy sediments, belong to the colder arctic waters. Such arctic faunæ are those of the Quebec group and of the Utica shale, and to some extent that of the Hamilton group.
Lyell’s theory on this subject has, I think, in some recent publications, been somewhat misapprehended. It is true that he stated hypothetically two contrasted conditions of distribution, in one of which all the land was equatorial, in another all polar; but he did not suppose that these conditions had actually occurred; and even in his earlier editions, before the recent discoveries and discussions as to ocean currents, he was always careful to attach due value to these in connection with subsidences and elevations.[EX] In his later editions he introduced more full references to current action, and also stated Croll’s theory, but still maintained the validity of his original conclusions.
[EX] See “Principles of Geology,” edition of 1840, chapter vii.
The sufficiency of this Lyellian theory to account for the facts, in so far as plants are concerned, may, I think, be inferred from the course of the isothermal lines at present. The south end of Greenland is on the latitude of Christiania in Norway on the one hand, and of Fort Liard in the Peace River region on the other; and while Greenland is clad in ice and snow, wheat and other grains, and the ordinary trees of temperate climates, grow at the latter places,[EY] It is evident, therefore, that only exceptionally unfavourable circumstances prevent the Greenland area from still possessing a temperate flora, and these unfavourable circumstances possibly tell even on the localities with which we have compared it. Further, the mouth of the McKenzie River is in the same latitude with Disco, near which are some of the most celebrated localities of fossil Cretaceous and Tertiary plants. Yet the mouth of the McKenzie River enjoys a much more favourable climate and has a much more abundant flora than Disco. If north Greenland were submerged, and low land reaching to the south terminated at Disco, and if from any cause either the cold currents of Baffin’s Bay were arrested, or additional warm water thrown into the North Atlantic by the Gulf Stream, there is nothing to prevent a mean temperature of 45° Fahr. from prevailing at Disco; and the estimate ordinarily formed of the requirements of its extinct floras is 50°,[EZ] which is probably above rather than below the actual temperature required.
[EY] See “Macoun’s Report,” “Geological Survey of Canada,” and Richardson’s “Boat Voyage.”
[EZ] Heer. See, also, papers by Prof. Haughton and by Gardner in “Nature” for 1878.
Since, then, geological facts assure us of mutations of the continents much greater than those apparently required to account for the changes of climate implied in the existence of the ancient arctic floras, it does not seem absolutely necessary to invoke any others.[FA] If, however, there are other true causes which might either aid or counteract those above referred to, it may be well to consider them.
[FA] Sir William Thomson, “Transactions of the Geological Society of Glasgow,” February 22, 1878.
Mr. Croll has, in his valuable work “Climate and Time” and in various memoirs, brought forward an ingenious astronomical theory to account for changes of climate. This theory, as stated by himself in a recent paper,[FB] is that when the eccentricity of the earth’s orbit is at a high value, and the northern winter solstice is in perihelion, agencies are brought into operation which make the southeast trade-winds stronger than the northeast, and compel them to blow over upon the northern hemisphere as far as the Tropic of Cancer. The result is that all the great equatorial currents of the ocean are impelled into the northern hemisphere, which thus, in consequence of the immense accumulation of warm water, has its temperature raised, so that ice and snow must to a great extent disappear from the arctic regions. In the prevalence of the converse conditions, the arctic zone becomes clad in ice, and the southern has its temperature raised.
[FB] “Cataclysmic Theories of Geological Climate,” “Geological Magazine,” May, 1878.