Lesquereux notices this species as characteristic of the lower part of the Carboniferous in Arkansas.
It will be observed that I regard the striated and ribbed stems not as internal axes, but as representing the outer surface of the plants. This was certainly the case with the present species and with C. Suckovii and C. nodosus. Other species, and especially those which belonged to Calamodendron, no doubt had a smooth or irregularly wrinkled external bark; but this gives no good ground for the manner in which some writers on this subject confound Calamites with Calamodendra, and both with Asterophyllites and Sphenophyllum. With this no one who has studied these plants, rooted in their native soils, and with their appendages still attached, can for a moment sympathise. One of the earliest geological studies of the writer was a bed of these erect Calamites, which he showed to Sir C. Lyell in 1844, and described in the “Proceedings of the Geological Society” in 1851, illustrating the habit of growth as actually seen well exposed in a sandstone cliff. Abundant opportunities of verifying the conclusions formed at that time have since occurred, the results of which have been summed up in the figures in Acadian Geology, which, though they have been treated by some botanists as merely restorations, are in reality representations of facts actually observed.
On these subjects, without entering into details, and referring for these to the elaborate discussions of Schimper, Williamson, and McNab, and to my paper on the subject, “Journal of the Geological Society,” vol. xxvii, p. 54, I may remark:
1. That the aërial stems of ordinary Calamites had a thin cortical layer, with lacunæ and fibrous bundles and multiporous vessels—the whole not differing much from the structure of modern Equiseta.
2. Certain arborescent forms, perhaps allied to the true Calamites, as well as possibly the old underground stems of ordinary species[DC] assumed a thick-walled character in which the tissues resembled the wedges of an exogen, and abundance of pseudo-scalariform fibres were developed, while the ribbing of the external surface became obsolete or was replaced by a mere irregular wrinkling.
[DC] Williamson, “Transactions of the Royal Society.” McNab, in “Proceedings of the Edinburgh Botanical Society.”
3. Sufficient discrimination has not been exercised in separating casts of the internal cavities of Calamites and Calamodendron from those representing other surfaces and the proper external surface.
4. There is no excuse for attributing to Calamites the foliage of Annularia, Asterophyllites, and Sphenophyllum, since these leaves have not been found attached to true Calamite stems, and since the structure of the stems of Asterophyllites as described by Williamson, and that of Sphenophyllum as described by the writer,[DD] are essentially different from those of Calamites.
[DD] “Journal of the Geological Society,” 1866.