The question of division lines of formations is one much agitated in the case of the Cambrian rocks. Whether certain beds are to be called Cambrian or Silurian has been a point greatly controverted; and the terms Primordial and Primordial Silurian have been used as means to avoid the raising of this difficulty. Many of our division lines in geology are arbitrary and conventional, and this may be the case with that between the Primordial and Silurian, the one age graduating into the other. There appears to be, however, the best reason to recognise a distinct Cambrian period, preceding the two great periods, those of the second and third faunas of Barrande, to which the term Silurian is usually applied. On the other hand, in so far as our knowledge extends at present, a strongly marked line of separation exists between the Laurentian and Primordial, the latter resting on the edges of the former, which seems then to have been as much altered as now. Still a break of this kind may be, perhaps must be, merely local; and may vary in amount. Thus, in some places we find rocks of Silurian and later ages resting directly on the Laurentian, without the intervention of the Primordial. In any case, where a line of coast is steadily sinking, each succeeding deposit will overlap that which went before; and this seems to have been the case with the Laurentian shore when the Primordial and Silurian were being deposited. Hence over large spaces the Primordial is absent, being probably buried up, except where exposed by denudation at the margin of the two formations.
This occurs in several parts of Canada, while the Laurentian rocks have evidently been subjected to metamorphism and long-continued weathering before the Lower Silurian were deposited; and in some cases the latter rest on weather-worn and pitted surfaces, and are filled with angular bits of the underlying rock, as well as with drift-shells which have been cast on these old Laurentian shores; while in other cases the Silurian rests on smooth water-worn Laurentian rocks, and is filled at the junction with well-rounded pebbles and grains of sand which have evidently been subjected to a more thorough attrition than those of the present beach. With respect to the line of division between the Primordial and the next succeeding rocks, it will be seen that important movements of the continents occurred at the close of the Cambrian, and in some places the Cambrian rocks have been much disturbed before the deposition of the Lower Silurian.
Seated on some ancient promontory of the Laurentian, and looking over the plain which, in the Primordial and Lower Silurian periods was the sea, I have often wished for some shred of vegetable matter to tell what lived on that land when the Primordial surf beat upon its shore, and washed up the Trilobites and Brachiopods of those old seas; but no rock has yet taken up its parable to reveal the secret, and the Primordial is vocal only with the old story: “And God said, Let the waters swarm with swarming living things, and it was so.” So our picture of the period may represent a sea-bottom swarming with animals of low grade, some sessile, some locomotive; and we may merely suppose a distant shore with vegetation dimly seen, and active volcanoes; but a shore on which no foot of naturalist has yet trod to scan its productions. Very different estimates have been formed of the amount of life in this period, according to the position given to its latest limit. Taking some of the more modern views of this subject, we might have included among the Primordial animals many additional creatures, which we prefer noticing in the Silurian, since it may at least be affirmed that their head-quarters were in that age, even if they had a beginning in the Primordial. It may be interesting here, however, to note the actual amount of life known to us in this period, taken in its largest scope. In doing this, I shall take advantage of an interesting table given by Dr. Bigsby,[G] and representing the state of knowledge in 1868, and shall group the species in such a manner as to indicate the relative abundance of distinct types of structure. We find then—
| Plants (all, or nearly all, supposed
to be sea-weeds, and some, probably, mere tracks or trails of animals) | 22 | species. |
| Sponges, and similar creatures | 27 | ” |
| Corals and their allies | 6 | ” |
| Starfishes and their allies | 4 | ” |
| Worms | 29 | ” |
| Trilobites and other crustaceans | 442 | ” |
| Lamp-shells and other molluscoids | 193 | ” |
| Common bivalve mollusks | 12 | ” |
| Common univalve mollusks and their allies | 172 | ” |
| Higher mollusks, nautili, cuttle-fishes, etc. | 65 | ” |
| In all | 972 | ” |
[G] “Thesaurus Siluricus.”
Now in this enumeration we observe, in the first place, a representation of all the lower or invertebrate groups of the waters. We have next the remarkable fact that the Radiata of Cuvier, the lowest and most plant-like of the marine animals, are comparatively slenderly represented, yet that there are examples of their higher as well as of their lower forms. We have the further fact that the crustaceans, the highest marine animals of the annulose type, are predominant in the waters; and that in the mollusks the highest and lowest groups are most plentiful, the middle less so. The whole number of species is small, and this may arise either from our having here reached an early period in the history of life, or from our information being defective. Both are probably true. Still, of the animals known, we cannot say that the proportions of the different kinds depend on defective knowledge. There is no reason, for example, why corals should not have been preserved as well as Trilobites, or why Brachiopods should have been presurved rather than ordinary bivalves. The proportions, therefore, it may be more safe to reason from than the aggregate. In looking at these proportions, and comparing them with those of modern seas, we are struck with the great number of species representing some types either now extinct or comparatively rare: the Trilobites and Brachiopods more particularly. We are astonished at the enormous preponderance of these two groups, and especially of the Trilobites. Further, we observe that while some forms, like Lingula and Nautilus, have persisted down to modern times, others, like the Trilobites and Orthids, perished very early. In all this we can dimly perceive a fitness of living things to physical conditions, a tendency to utilise each type to the limit of its capacities for modification, and then to abandon it for something higher; a tendency of low types to appear first, but to appear in their highest perfection and variety; a sudden apparition of totally diverse plans of structure subserving similar ends simultaneously with each other, as for instance those of the Mollusk and the Crustacean; the appearance of optical and mechanical contrivances, as for example the compound eyes of the Trilobite and the swimming float of the Orthoceras, in all their perfection at first, just as they continue to this day in creatures of similar grade. That these and other similar things point to a uniform and far-reaching plan, no rational mind can doubt; and if the world had stopped short in the Primordial period, and attained to no further development, this would have been abundantly apparent; though it shines forth more and more conspicuously in each succeeding page of the stony record. How far such unity and diversity can be explained by the modern philosophy of a necessary and material evolution out of mere death and physical forces, and how far it requires the intervention of a Creative mind, are questions which we may well leave with the thoughtful reader, till we have traced this history somewhat further.