15. The fluids, composed principally of water holding solid matter in solution, or in a state of mechanical division, either contribute to the formation of the blood, or constitute the blood, or are derived from the blood; and after having served some special office in a particular part of the system, are returned to the blood; and according to the nature and proportion of the substances they contain, are either aqueous, albuminous, mucous, gelatinous, fibrinous, oleaginous, resinous, or saline.

16. When the analysis of the different kinds of animal matter that enter into the composition of the body has been carried to its ultimate point, it appears to be resolvable into two primitive forms: first, a substance capable of coagulation, but possessing no determinate figure; and secondly, a substance having a determinate figure and consisting of rounded particles. The coagulable substance is capable of existing by itself; the rounded particles are never found alone, but are invariably combined with coagulated or coagulable matter. Alone or combined with the rounded particles, the coagulable matter forms, when liquid, the fluids, when coagulated, the solids.

17. When solid, the coagulable substance is disposed in one of two forms, either in that of minute threads or fibres, or in that of minute plates or laminæ; hence every solid of the body is said to be either fibrous or laminated. The fibres or laminæ are variously interwoven and interlaced, so as to form a net-work or mesh; and the interspaces between the fibres or laminæ are commonly denominated areolæ or cells (fig. XVII).

18. This concrete substance, fibrous or laminated, is variously modified either alone or in combination with the rounded particles. These different modifications and combinations constitute different kinds of organic substance. When so distinct as obviously to possess a peculiar structure and peculiar properties, each of these modifications is considered as a separate form of organized matter, and is called a PRIMARY TISSUE. Anatomists and physiologists have been at great pains to discriminate and classify these primary tissues; for it is found that when employed in the composition of the body, each preserves its peculiar structure and properties wherever placed, however combined, and to whatever purpose applied, undergoing only such modification as its local connexions and specific uses render indispensable. Considering every substance employed in the construction of the body, not very obviously alike, as a distinct form of organized matter, these primary tissues may be said to consist of five, namely, the membranous, the cartilaginous, the osseous, the muscular, and the nervous.

19. The first primary tissue is the peculiar substance termed MEMBRANE. It has been already stated (16) that one of the ultimate forms of animal matter is a coagulable substance, becoming concrete or solid under the process of coagulation. The commencement of organization seems to be the arrangement of this concrete matter into straight thready lines, at first so small as to be imperceptible to the naked eye. Vast numbers of these threads successively uniting, at length form a single thread of sufficient magnitude to be visible, but still smaller than the finest thread of the silkworm. If the length of these threads be greater than their breadth, they are called fibres; if, on the contrary, their breadth exceed their length, they are termed plates or laminæ. By the approximation of these fibres or plates in every possible direction, and by their accumulation, combination, and condensation, is constituted the simplest form of organized substance, the primary tissue called membrane.

20. Membrane once formed is extensively employed in the composition of the body: it is indeed the material principally used in producing, covering, containing, protecting, and fixing every other component part of it. It forms the main bulk of the cartilaginous tissue; it receives into its cells the earthy matter on which depend the strength and hardness of the osseous tissue; it composes the canals or sheaths in which are deposited the delicate substance of the muscular, and the still more tender pulp of the nervous tissue; it gives an external covering to the entire body; it lines all its internal surfaces; it envelopes all internal organs; it enters largely as a component element into the substance of every organ of every kind; it almost wholly constitutes all the internal pouches and sacs, such as the stomach, the intestines, the bladder; and all tubes and vessels, such as arteries, veins, and lymphatics; it furnishes the common substance in which all the parts of the body are, as it were, packed; it fills up the interstices between them; it fixes them in their several situations; it connects them all together; in a word, it forms the basis upon which the other parts are superinduced; or rather the mould into which their particles are deposited; so that were it possible to remove every other kind of matter, and to leave this primary tissue unaltered in figure and undiminished in bulk, the general form and outline of the body, as well as the form and outline of all its individual parts, would remain unchanged.

21. The properties which belong to membrane are cohesion, flexibility, extensibility, and elasticity. By its property of cohesion, the several parts of the body are held together; by its combined properties of cohesion, flexibility, and extensibility, the body in general is rendered strong, light, and yielding, while particular parts of it are made capable of free motion. But elasticity, that property by which parts removed from their situation in the necessary actions of life are restored to their natural position, may be regarded as its specific property. The varied purposes accomplished in the economy by the property of elasticity will be apparent as we advance in our subject. Meantime, it will suffice to observe that it is indispensable to the action of the artery in the function of the circulation; to the action of the thorax in the function of respiration; to the action of the joints in the function of locomotion: in a word, to the working of the entire mechanism by which motion of every kind and degree is effected. All these properties are physical, not vital; vital properties do belong even to this primary form of animal matter; but they are comparatively obscure. In the tissue with which organization commences, and which is the least removed from an inorganic substance, the properties that are prominent and essential are merely physical.

22. By chemical analysis, membrane is found to contain but a small proportion of azote, the peculiar element of animal matter. Its proximate principles are gelatin, albumen, and mucus. In infancy and youth, gelatin is the most abundant ingredient; at a more advanced period, albumen predominates[3]. Gelatin differs from albumen in containing a less proportion of azote and a greater proportion of oxygen; on both accounts it must be regarded as less animalized. Thus animalization bears a certain relation to organization. The simplest animal tissue is the least animalized, and the least of all at the earliest period of life. Not only are the physical and mental powers less developed in the young than in the adult, but the very chemical composition of the primary tissue of which the body is constructed is less characteristic of the perfect animal.

23. Membrane exists under several distinct forms; a knowledge of the peculiarities of which will materially assist us in understanding the composition of the body. The simplest form of membrane, and that which is conceived to constitute the original structure from which all the others art produced, is termed the cellular. When in thin slices, cellular membrane appears as a semi-transparent and colourless substance; when examined in thicker masses, it is of a whitish or greyish colour. It consists of minute threads, which cross each other in every possible direction, leaving spaces between them, and thus forming a mesh or net-work (fig. XVII.), not unlike the spider's web. The term cells, given to these interspaces, is employed rather in a figurative sense than as the expression of the fact; for there are no such distinct partitions as the term cell implies. The best conception that can be formed of the arrangement of the component parts of this structure is, to suppose a substance consisting of an infinite number of slender thready lines crossing each other in every possible direction (fig. XVII.). The interspaces between these lines during life, and in the state of health, are filled with a thin exhalation of an aqueous nature, a vapour rather than a fluid, rendering and keeping the tissue always moist. This vapour consists of the thinner part of the blood, poured into these interstitial spaces by a process hereafter to be described, termed secretion. When occupying those spaces, it makes no long abode within them, but is speedily removed by the process of absorption. In health, these two operations exactly equal each other; but if any cause arise to disturb the equilibrium, the vapour accumulates, condenses and forms an aqueous fluid, which distends the cells and gravitates to the most depending parts. Slightly organized as this tissue is, and indistinct as its vita functions may be, it is obvious that it must be the seat of at least two vital functions, secretion and absorption.