1. The muscle called the Scalenus. 2. The muscles called the Intercostals. 3. Subclavius. 4. The bone called the Clavicle. 5. The muscle called the Serratus Magnus Anticus. 6. Obliquius Externus. 7. Rectus. 8. Obliquius Internus.
391. The first step necessary to the ascent of the diaphragm is the relaxation of its muscular fibres. As soon as these fibres are in a state of relaxation, that is, when the organ has changed from an active to a completely passive state, the powerful muscles of the abdomen (fig. [CXLVI]. 6, 7, 8) contract, and push the abdominal viscera and the diaphragm with them upwards towards the cavity of the chest (fig. [CXLV]. 2); and thus, by the descent of the ribs and the ascent of the diaphragm, the capacity of the thorax is diminished in every direction, and the motion of expiration is completed.
392. Such is the mechanism by which the capacity of the thorax is alternately enlarged and diminished in the two alternate states of inspiration and expiration, and the mechanism thus adjusted works in the following mode.
393. Expiration succeeding to the state of inspiration, the ribs descend, the diaphragm ascends, the capacity of the thorax lessens, and the compressed lungs are forced within the smallest possible space. Then, inspiration, succeeding to the state of expiration, the ribs ascend and the diaphragm descends; the capacity of the thorax is enlarged, and the lungs freed from their pressure expand and fill the greater space obtained. In about a second and a half after the state of inspiration has been induced, that of expiration recommences; the motion of inspiration occupying about double the time of the motion of expiration, and these alternate conditions succeed each other in a regular and uniform course, day and night, during our sleeping and our waking hours to the end of life.
394. As long as the function is performed in a perfectly natural manner, a given number of these alternate movements takes place in a certain time, constituting what is termed the rhythm of the respiratory motions. These motions perfectly regular in number and time, are likewise, in the natural state of the function, performed only with a certain degree of energy; but they are variously modified at the command of the will; in obedience to numerous sensations and emotions; in the performance of a great variety of complex actions, and in different states of disease. These modifying circumstances may cause the action of inspiration to be more full and deep, and that of expiration to be more forcible and complete than natural; or they may cause both movements to be shorter and quicker than common: hence the distinction of respiration into ordinary and extraordinary.
395. In ordinary respiration, that is, when the respiratory motions are perfectly calm and easy, the ascent and descent of the ribs are scarcely perceptible; the action is confined almost exclusively to the ascent and descent of the diaphragm. In this condition the only action of the intercostal muscles is to fix the ribs, and thus to afford one of the two fixed points which have been shown ([385]) to be essential to the action of the diaphragm. But in extraordinary respiration, that is, when circumstances happen in the economy which require that those motions should be extended, auxiliary sources can be put in requisition. There are many powerful muscles situated about the breast, shoulder and back (fig. [CXLVI]. and [CXLVII]. ); which are capable of elevating the ribs and protruding the sternum to a very considerable extent (figs. [CXLVI]. 1, 2, 3, 5; and [CXLVII]. 1, 2, 3). Where, for example, the fullest inspiration which it is possible to take is required, the bones of the shoulder and shoulder-joint are firmly fixed by resting the hands upon the knees, and then every muscle which has the slightest connexion with the thorax, either before or behind, capable of raising the ribs, is added to the inspiratory apparatus (figs. CXLIV. and CXLVII.); at the same time that the abdominal muscles are relaxed to the utmost degree, in order to facilitate the ascent of the ribs and the descent of the diaphragm (figs. [CXLIV]. 2, and [CXLVI]. 6, 7, 8). If, on the contrary, the fullest possible expiration is required, the abdominal muscles contract most forcibly (fig. [CXLV]. 2), and every other muscle which is capable of still farther depressing the ribs and of elevating the diaphragm (fig. [CXLVI]. 6, 7, 8) is called into intense action. By these forcible and extraordinary efforts the thorax may be enlarged or diminished double its ordinary capacity.
Fig. CXLVII.—View of Muscles which are capable of assisting in elevating the Ribs and protruding the Sternum, in states of extraordinary respiration.
1. The muscle called the Great Pectoral. 2. The Small Pectoral. 3. The Serratus Magnus.
396. Such are the mechanism and action of the powers which communicate to the thorax, the motions by which its capacity is alternately enlarged and diminished, and by which the requisite impulse is communicated to the fluids which flow to and from the lungs in the different states of respiration; that is, by which air and blood flow to the lungs in the action of inspiration, and from the lungs in the action of expiration.