581. In man, the several classes of the teeth are so similarly developed, so perfectly equalized, and so identically constructed, that they may be considered as the true type from which all the other forms are deviations.

582. For the accomplishment of their office the teeth must be endowed with prodigious strength: for the fulfilment of purposes immediately connected with the apparatus of digestion, it is necessary that they should be placed in the neighbourhood of exceedingly soft, delicate, irritable, and sentient organs. That they may possess the requisite degree of strength, they are constructed chiefly of bone, the hardest organized substance. Bone, though not as sensible as some other parts of the body, is nevertheless sentient. The employment of a sensitive body in the office of breaking down the hard substances used as food would be to change the act of eating from a pleasurable into a painful operation. It has been shown (vol. i. p. 84) that provision is made for supplying to the animal a never-failing source of enjoyment in the annexation of pleasurable sensations with the act of eating, and that, taking the whole of life into account, the sum of enjoyment secured by this provision is incalculable. But all this enjoyment might have been lost, might even have been changed into positive pain, nay, must have been changed into pain, but for adjustments numerous, minute, delicate, and, at first view, incompatible.

583. Had a highly-organized and sensitive body been made the instrument of cutting, tearing, and breaking down the food, every tooth, every time it comes in contact with the food, would produce the exquisite pain now occasionally experienced when a tooth is inflamed. Yet a body wholly inorganic and therefore insensible, could not perform the office of the instrument; first, because a dead body cannot be placed in contact with living parts without producing irritation, disease, and consequently pain; and, secondly, because such a body being incapable of any process of nutrition, must speedily be worn away by friction, and there could be no possibility of repairing or of replacing it. The instrument in question, then, must possess hardness, durability, and, to a certain extent, insensibility; yet it must be capable of forming an intimate union with sentient and vital organs, must be capable of becoming a constituent part of the living system.

584. To communicate to it the requisite degree of hardness, the hard substance forming its basis is rendered so much harder than common bone that some physiologists have even doubted whether it be bone, whether it really possess a true organic structure. That there is no ground for such doubt the evidence is complete. For,

1. The tooth, like bone in general, is composed partly of an earthy and partly of an animal substance; the earthy part being completely removable by maceration in an acid, and the animal portion by incineration, the tooth under each process retaining exactly its original form.

2. The root of the tooth is covered externally by periosteum; its internal cavity is lined by a vascular and nervous membrane, and both structures are intimately connected with the substance of the tooth. If these membranes really distribute their blood-vessels and nerves to the substance of the tooth, which there is no reason to doubt, the analogy is identical between the structure of the teeth and that of bone.

3. Though the blood-vessels of the teeth are so minute that they do not, under ordinary circumstances, admit the red particles of the blood, and though no colouring matter hitherto employed in artificial injections has been able, on account of its grossness, to penetrate the dental vessels, yet disease sometimes accomplishes what art is incapable of effecting. In jaundice the bony substance of the teeth is occasionally tinged with a bright yellow colour; and in persons who have perished by a violent death, in whom the circulation has been suddenly arrested, it is of a deep red colour. Moreover, when the dentist files a tooth, no pain is produced until the file reaches the bony substance; but the instant it begins to act upon this part of the tooth, the sensation becomes sufficiently acute.

585. These facts demonstrate that the bony matter of the tooth, though modified to fit the instrument for its office, is still a true and proper organized substance.

586. Each tooth is divided into body, neck, and root (fig. [CLVIII]. 1, 2, 3). The body is that part of the tooth which is above the gum, the root that part which is below the gum, and the neck that part where the body and the root unite (fig. [CLVIII]. ). The body, the essential part, is the tooth properly so called, the part which performs the whole work for which the instrument is constructed, to the production and support of which all the other parts are subservient.

Fig. CLVIII.