601. As long as the operations of mastication and insalivation go on, the mouth forms a closed cavity from which the food cannot escape; for the lips enclose it before, the cheeks at the sides, the tongue below, and the soft palate behind, the inferior edge of which being applied in close and firm contact with the base of the tongue, prevents all communication between the mouth and the pharynx.
602. When, by mastication, the food is sufficiently divided, and by insalivation softened and animalized to fit it for the future changes it is to undergo, it is collected by the tongue, and carried by that organ to the back part of the mouth. The soft palate (fig. CLII. 1), obedient to the stimulus of the duly prepared food, rises the instant it is touched by it, and affords it a free passage to the pharynx (figs. [CLIII]. . 10, and [CLIV]. . 10).
603. The pharynx (fig. [CLIII]. 10), a muscular bag, immediately continuous with the mouth (fig. [CLIII]. 1), is a vestibule into which open several highly important organs. Before is the entrance to the windpipe, termed the glottis (fig. [CLIV]. 9), leading directly to the larynx (fig. [CLIV]. 8); at the sides are the mouths of two ducts, termed the Eustachian tubes, which lead to the internal part of the organ of hearing; above are two entrances to the nose (fig. [CLIV]. 1); and below is the passage to the stomach (fig. [CLIII]. 12).
604. Were the food to enter the Eustachian tubes or the nose, it would occasion great inconvenience; were it to enter the glottis, it would cause death. It is prevented from entering the Eustachian tubes and the nose by the soft palate (fig. CLII. 1 and 2), which by the very act of rising to afford an opening from the mouth to the pharynx, is carried over the other apertures so as completely to close them. By the varied direction of the muscular fibres which enter into the composition of this organ, it is enabled to execute the different and even opposite motions required in the performance of its important office.
605. The food is prevented from entering the glottis partly by a cartilaginous valve (fig. [CLIV]. 7), termed the epiglottis, placed immediately above the glottis, and attached to the root of the tongue (fig. [CLIV]. 6). In delivering the food to the pharynx the tongue passes backwards (fig. [CLIV]. 6). In passing backwards it pushes in the same direction the epiglottis which is attached to it, and so necessarily carries it over the glottis, completely closing the aperture (fig. [CLIV]. 9). At the same time the opening is still more securely closed by the glottis itself, in consequence of the powerful and simultaneous contraction of the muscles that act upon it in the production of the voice. It is proved, by direct experiment, that the spontaneous closure of the glottis is a more powerful agent in excluding the food from the larynx even than the depression of the epiglottis; but both organs concur in producing the same result; and a double security is provided against an event which would be fatal.
606. It is deeply interesting to observe the part performed in these operations by sensation and volition, and the boundary at which their influence terminates and consciousness itself is lost. Mastication, a voluntary operation, carried on by voluntary muscles, at the command of the will, is attended with consciousness, always in the state of health of a pleasurable nature. To communicate this consciousness, the tongue, the palate, the lips, the cheeks, the soft palate, and even the pharynx, are supplied with a prodigious number of sentient nerves. The tongue especially, one of the most active agents in the operation, is supplied with no less than six nerves derived from three different sources. These nerves, spread out upon this organ, give to its upper surface a complete covering, and some of them terminate in sentient extremities visible to the naked eye. These sentient extremities, with which every point of the upper surface, but more especially the apex, is studded, constitute the bodies termed papillæ, the immediate and special seat of the sense of taste. This sense is also diffused, though in a less exquisite degree, over the whole internal surface of the mouth. Close to the sense of taste is placed the seat of the kindred sense of smell. The business of both these senses is with the qualities of the food. Mastication at once brings out the qualities of the food and puts the food in contact with the organs that are to take cognizance of it. Mastication, a rough operation, capable of being accomplished only by powerful instruments which act with force, is carried on in the very same spot with sensation, an exquisitely delicate operation, having its seat in soft and tender structures, with which the appropriate objects are brought into contact only with the gentlest impulse. The agents of the coarse and the delicate, the forcible and the gentle operations are in close contact, yet they work together not only without obstruction, but with the most perfect subserviency and co-operation.
607. The movements of mastication are produced, and, until they have accomplished the objects of the operation, are repeated by successive acts of volition. To induce these acts, grateful sensations are excited by the contact of the food with the sentient nerves so liberally distributed over almost the whole of the apparatus. To the provision thus made for the production of pleasurable sensation, is superadded the necessity of direct and constant attention to the pleasure included in the gratification of the taste. It is justly observed by Dr. A. Combe, that without some degree of attention to the process of eating, and some distinct perception of its gratefulness, the food cannot be duly digested. When the mind is so absorbed as to be wholly unconscious of it, or even indifferent to it, the food is swallowed without mastication; then it lies in the stomach for hours together without being acted upon by the gastric juice, and if this be done often, the stomach becomes so much disordered as to lose its power of digestion, and death is the inevitable result: so that not only is pleasurable sensation annexed to the reception of food, but the direct and continuous consciousness of that pleasurable sensation during the act of eating is made one of the conditions of the due performance of the digestive function.
608. With the operation of mastication and one part of the process of deglutition, immediately to be noticed, the agency of volition and sensation cease. Beyond this the function of digestion is wholly an organic process. In addition to the reasons assigned (vol. i. p. 55) why all the organic processes are placed alike beyond the cognizance of sense and the control of the will, there is this special reason why, in the function of digestion, they cease at the exact boundary assigned them.
609. Every time the act of deglutition is performed the openings to the windpipe and to the nose are closed, so that during this operation all access of air to the lungs is stopped, consequently it is necessary that the passage of the food through the pharynx should be rapid. Mastication, a voluntary process, may be performed slowly or rapidly, perfectly or imperfectly, without serious mischief; but life depends on the passage of the food through the pharynx with extreme rapidity and with the nicest precision. It is therefore taken out of the province of volition and entrusted to organs which belong to the organic life, organs which carry on their operations with the steadiness, constancy, and exactness of bodies whose motions are determined by a physical law.
610. No sooner does the duly-prepared food touch the soft palate than the whole apparatus of deglutition is instantly in motion. This movable partition suddenly rises to afford to the food a free passage to the pharynx. The pharynx itself, at the same instant, rises to receive the morsel thrust towards it by the pressure of the tongue; and one muscle, the stylo-pharyngeus, which concurs in producing this movement, seems specially intended, in addition, to expand the pharynx. Three muscles throw their fibres around the pharynx, termed its upper, middle, and lower constrictors, which, the moment the morsel reaches the pharynx, contract upon it, and embrace it firmly. At the same instant the larynx, closing its aperture, springs forward towards the base of the tongue, under which it is in a manner concealed, the additional shield of the epiglottis being simultaneously thrown over the glottis. By this movement of the larynx, upwards and forwards, the course of the morsel across the dangerous passage is shortened. All these motions take place with such rapidity that Boerhaave said the action is convulsive. And now the food, firmly pressed by the pharynx, cannot return to the mouth, for the root of the tongue is there stopping up the passage; it cannot enter the Eustachian tubes or the nose, for the soft palate is there closing the apertures; it cannot enter the larynx, for a double guard is placed upon the glottis securing its firm closure. The food can advance in one direction only, the direction required, that which leads to the esophagus. Well, therefore, on the contemplation of these complex structures and the consent and harmony with which they act, might Paley say, “In no apparatus put together by art do I know such multifarious uses so aptly contrived as in the natural organization of the human mouth and its appendages. In this small cavity we have teeth of different shape; first, for cutting; secondly, for grinding; muscles most artificially disposed for carrying on the compound motions of the lower jaw by which the mill is worked; fountains of saliva springing up in different parts of the cavity for the moistening of the food while the mastication is going on; glands to feed the fountains; a muscular contrivance in the back part of the cavity for the guiding of the prepared aliment into its passage towards the stomach, and for carrying it along that passage. In the mean time, and within the same cavity, is going on other business wholly different, that of respiration and of speech. In addition, therefore, to all that has been mentioned, we have a passage opened from this same cavity of the mouth into the lungs for the admission of air, for the admission of air exclusively of every other substance; we have muscles, some in the larynx, and, without number, in the tongue, for the purpose of modulating that air in its passage, with a variety, a compass, and a precision of which no other musical instrument is capable; and, lastly, we have a specific contrivance for dividing the pneumatic part from the mechanical, and for preventing one set of functions from interfering with the other. The mouth, with all these intentions to serve, is a single cavity; is one machine, with its parts neither crowded nor confined, and each unembarrassed by the rest.” It should be added, the mouth is also the immediate seat of one of the senses, and is in intimate communication with a second sense; both these senses are always excited while the principal business performed by the machine is carried on, and are necessarily excited by the very working of the machine, and the sensations induced in the natural and sound state of the apparatus are invariably pleasurable.