1. A solvent power. The first action of the stomach on the alimentary substances presented to it is to reduce them to a fluid state. No substance is nutritious which is not a fluid, or capable of being reduced to a fluid. The stomach reduces alimentary substances to a fluid state by combining them with water. Water enters into the composition of organized bodies in two states, as an essential and as an accidental element. A quantity of water is contained in sugar when reduced to its dryest state; this water cannot be dissipated without the decomposition of the sugar; it is therefore an essential constituent of the compound. Water is combined with sugar in its moist state: of this water much may be removed without destroying the essential properties of the sugar: this part of the water is therefore said to be an accidental constituent of the sugar. In most cases organized bodies contain water in both these forms; and though it is commonly impossible to discriminate between the water that is essential and that which is accidental, yet the mode of union among the elements of bodies in these two states of their combination with water are essentially different. The stomach has the power of combining water with alimentary substances in both these forms. Thus fluid albumen, or white of egg, presented to the stomach is immediately coagulated or converted into a solid. Soon this solid begins to be softened, and the softening goes on until it is again reduced to a fluid. What was fluid albumen in the white of egg is now fluid albumen in chyme; but the albumen has undergone a remarkable change. Out of the stomach the albumen of the egg may be converted by heat into a firm solid; but the albumen of the chyme is capable of being converted only into a loose and tender solid. In passing from its state in the egg to its state in the chyme, the albumen has combined with a portion of water which has entered as an essential ingredient into its composition. By this combination the compound is reduced from what may be called a strong to a weak state. This is the first action exerted by the stomach on most alimentary substances. They are changed from a concentrated to a diluted, from a strong to a weak state: the power by which the stomach effects this change is called its reducing power, and the agent by which it accomplishes it is the gastric juice; the essential ingredient of which has been shown to be muriatic acid, or chlorine (639, et seq.). The muriatic acid obtained from the common salt of the blood is poured in the form of gastric juice into the stomach, dissolves the food, combines it with water, reduces it from a concentrated solid to a dilute fluid; and thus brings it into the condition proper for the subsequent part of the process.
2. A converting power. Since whatever be the varieties of food, the chyme invariably forms a homogeneous fluid, the stomach must be endowed with the power of transforming the simple alimentary principles into one another; the saccharine into the oily, and the oily into the albuminous. The transformation of the saccharine into the oleaginous principle is traceable out of the body in the conversion of sugar into alcohol, which is essentially an oil. That the same transformation takes place within the body is indubitable. The oleagenous and the albuminous principles are already so nearly allied in nature to animal substance that they do not need to undergo any essential change in their composition.
3. A completing power. When the alimentary substances have been reduced and formed into chyme, when the chyme has been converted into chyle, and when the chyle absorbed by the lacteals is transmitted to the mesenteric glands, it undergoes during its passage through these organs a process the direct reverse of that to which it is subjected in the stomach; for whereas it is the office of the stomach to combine the alimentary substances with water, it is one office of the mesenteric glands to remove the superfluous water of the chyle; to abstract whatever particles of matter may be contained in the compound which are not indispensable to it, and to concentrate its essential constituents; and consequently these organs exert on the digested aliment a completing, in contradistinction to a reducing power.
4. A vitalizing power. When sugar is converted into oil, when oil is converted into albumen, when albumen, by the successive processes to which it is subjected is completed, that is, when the alimentary substances are made to approximate in the closest possible degree to the nature of animal substance, they must undergo a still further change, more wonderful than any of the preceding, and far more inscrutible; they must be endowed with vitality; must be changed from dead into living matter. Living substance only is capable of forming a constituent part of living substance. The ultimate action of the digestive organs is the communication of life to the food, to which last and crowning process the reducing, converting, and completing processes are merely subordinate and preparatory. Of the agency by which this process is effected we are wholly ignorant; we know that it goes on; but the mode in which it is accomplished is veiled in inscrutable darkness.
704. Blood is alive; blood is formed from the food; life is communicated to the food before it is mixed with the blood. The blood is essentially albumen, which it contains in the form of albumen properly so called, in that of fibrin, and in that of red particles. In the thoracic duct the strong albumen of the lymph is mixed with the weaker albumen of the chyle. At the point where the thoracic duct terminates in the venous system, lymph and chyle are mixed with venous blood, and all commingled are borne directly to the lungs. There the carbon with which the venous blood is loaded is expelled in the form of carbonic acid gas; the particles of the lymph undergo some, as yet, unknown change, exalting their organization; and the water hitherto held in chemical union with the weak albumen of the chyle, is separated and carried out of the system together with the carbonic acid gas in the form of aqueous vapour. By this removal of its aqueous particles the ultimate completion is given to the digested aliment; and the weak and delicate albumen of the chyle is converted into the strong and firm albumen of the blood.
705. It has been stated ([539]), that though gelatin enters abundantly into the composition of many tissues of the body, and performs most important uses in the economy, it is never found in the blood; that it is formed from the albumen of the blood by a reducing process, in consequence of which carbon is evolved, which unites with the free oxygen of the blood, forming carbonic acid, thus conducing, among other purposes, to the production of animal heat. It is equally remarkable, that though the lymphatics or absorbents arise in countless numbers from every tissue of the body, and are endowed with the power of taking up every constituent particle of every organ, solid as well as fluid, yet gelatin is never found in the lymphatic vessels. The lymphatics contain only albumen in a form far more proximate to the blood than that of the chyle; consequently, before the gelatin of the body is taken up by the lymphatics, it must be reconverted into albumen; that is, the absorbed gelatin must undergo a process analogous to that which gelatin and other matters undergo in the stomach and duodenum; it follows that the digestive process is not confined to the stomach and duodenum, but is carried on at every point of the body. Hence there are two processes of digestion, a crude and a refined process. The crude process is carried on in the stomach and duodenum, in which dead animal matter is converted into living substance, as yet, however, possessing only the lowest kind of vitality. The capillary arteries receiving the substance thus prepared for them, build it up into structure perhaps the lowest and coarsest, the least organized, and capable of performing only the inferior functions.
706. Capillary arteries in countless numbers terminate in the tissues in membraneless canals (304 and 310). Particles of the blood are seen to quit the arterial stream and to enter into the tissues, becoming a component part of them: other particles are seen to quit the tissues and to enter the current of the blood. The latter are probably organic particles, to which a certain degree of elaboration has been already given, now transmitted to the capillary veins, to be carried back to the lungs to undergo there a further depuration, fitting them on their return to the system for a higher organization.
707. Thus the lymphatic vessels, analogous in so many other respects to the veins, are probably similar to them in this also—that they take up from the tissues particles already organized, in order to submit them to processes which communicate to them a progressively higher organization. The notion that the contents of the lymphatics consist of worn-out particles, capable of accomplishing no further purpose in the economy, is not tenable:—
1. Because it is not analogous to the ordinary operations of nature to mix wholly excrementitious matter with a substance for the production, elaboration, and perfection of which, she has constructed such an expensive apparatus.
2. Because, on the other hand, the admixture of matter already highly animalized with matter, as yet but imperfectly animalized, exalts the nature of the latter, and is conducive to its complete animalization.