844. The various matters contained in organized bodies, and even those which enter as constituent elements into their composition, are constantly removed from the system, and thrown off into the external world. The matters thus rejected are called excretions; and the various processes by which their elimination is effected constitute a common function termed excretion.
845. Excretion is the necessary consequence of the deterioration which all organized matter undergoes by the actions of life. The matters removed by the process consist of the waste particles of the body, or the particles expended in the vital actions, as the aliment contains the particles which replenish the waste, and compensate the expenditure.
846. The excretions are separated from the common organized mass by processes perfectly analogous to those comprehended in the great function of secretion. Excretion is only a particular form of secretion: the difference between the two functions is, that, in the former, the matter eliminated being either noxious or useless, is separated for the sole purpose of being rejected; while, in the latter, the matter eliminated is destined to perform some useful purpose in the economy. Accordingly, the products of excretion are termed excrementitious; and those of secretion, recrementitious.
847. The chief matters excreted by the plant are oxygen, carbonic acid, air; water, in some few cases, under peculiar circumstances, ammonia and chlorine; and in still rarer cases, during the night, poisonous substances, as carburetted hydrogen, together with acrid, and even narcotic principles.
848. The forms under which these excretions are eliminated are exceedingly various. Sometimes the matter excreted is in the shape of gas, at other times it is in that of vapour, and at others in that of liquid. The chief gaseous exhalations are oxygen and carbonic acid; the vaporous exhalations consist principally of water, in the state of vapour; and the liquid exhalations are either pure water, or water holding in combination sugar, mucilage, and other proximate vegetable principles. Even the peculiar products formed by the vital actions of the plant, as the volatile oils, the fixed oils, the balsams, the resins, and perhaps, with the exception of gum, sugar, starch, and lignine, all the substances formed out of the proper juices of the plant, are true excretions; for these substances are fixed immovably in the cells, sacs, or tubes which secrete and contain them: they are not consumed in the growth of the plant; they do not appear to be applied to any useful purpose in the economy; they are injurious, and even poisonous to the very plant in which they are formed when taken up by the roots and combined with the sap: as long as they remain in the plant they are isolated in the individual parts in which they are first deposited, until with the advancing age of the plant they lose their aqueous particles, and are finally dried up; they, therefore, possess all the essential characters of excrementitious substances.
849. The organs by which these matters are excreted are the leaves, the flowers, the fruits, the roots, and certain bodies called glands.
850. The gaseous and vaporous exhalations are effected chiefly by the leaves, which it has been shown (320 and 465), under the influence of the solar ray, are always pouring out a large quantity of oxygen, and still larger quantities of fluid in the state of vapour.
851. Similar matters are exhaled by the flowers either in the form of vapour or of liquid; and this exhalation commonly bears with it a peculiar odour, which proceeds from an essential oil, sometimes evaporated with the pollen, and at other times secreted by glandular bodies which have their seat in the petals.
852. Fruits, and especially green fruits, as raspberries, pears, apples, plums, apricots, figs, cherries, gooseberries, and grapes, pour out oxygen during the day, and carbonic acid gas during the night, and thus co-operate with leaves in carrying on the function of excretion.
853. The more elaborate excretions contained in special receptacles, and formed by diverse organs from the proper juices of the plant, descend chiefly by the bark, and are poured by the roots into the soil. These excretions, if re-absorbed by the roots, and re-introduced into the system of the plant that has rejected them, poison that plant. Consequently, two processes of deterioration are always going on in the soil; first, the absorption of the nutrient matter contained in it; and, secondly, the accumulation of excrementitious matter constantly poured into it by the growing plant. By the addition of manure, the soil is replenished with fresh nutritive materials; by a rotation of crops, it is purified from noxious excretions. It is a remarkable and beautiful adjustment, that excrementitious substances which are destructive to plants of one natural family, actually promote the growth of plants of a different species. Thus, if wheat be sown upon a tract of land proper for that grain, it may produce a good crop the first, the second, and perhaps even the third year, as long as the ground is what the farmers call in good heart. But, after a time, it will yield no more of that particular kind of corn. Barley it may still bear, and, after this, oats, and perhaps after these, pease, or some other species belonging to a different family. The excrementitious matter deposited in the soil by a preceding is absorbed by a succeeding crop; the matter excreted by the former serving as nutriment or stimulus to the latter. But though in this mode all noxious matter is removed from the soil, yet the ground at last becomes quite barren, in consequence of having parted with all its nutrient particles, and then it will yield no more produce until it is supplied with a new fund of matter. This new matter is afforded by vegetable or animal substances, in which, the principle of life having become extinct, the peculiar bond that held their particles together is dissolved. Leaves, flowers, fruits, bark, roots; hair, skin, horns, hoofs, fat, muscle, bone, the blood itself, whatever has formed a part of the organized body, now dead, and repassing through the process of decomposition, back to the simple physical elements, all its forms of beauty gone, and exhaling only matters highly deleterious to animal life, mixed with the soil, are recombined into new products, spring up into new plants, and thus re-appear under new forms of beauty, and afford fresh nutriment to myriads of animals. The very refuse of the matters which have served as food and clothing to the inhabitants of the crowded city, and which, allowed to accumulate there, taint the air, and render it pestilential, promptly removed, and spread out on the surface of the surrounding country, give it healthfulness, clothe it with verdure, and endow it with inexhaustible fertility.