The pedicellariæ of the echinus have been already spoken of, and the difficulty as to their origin from minute, fortuitous, indefinite variations has been stated. But structures essentially similar (called avicularia, or "bird's-head processes") are developed from the surface of the compound masses of certain of the highest of the polyp-like animals (viz. the Polyzoa or, as they are sometimes called, the Bryozoa).
These compound animals have scattered over the surface of their bodies minute processes, each of which is like the head of a bird, with an upper and lower beak, the whole supported on a slender neck. The beak opens and shuts at intervals, like the jaws of the pedicellariæ of the echinus, and there is altogether, in general principle, a remarkable similarity between the structures. Yet the echinus can have, at the best, none but the most distant genetic relationship with the Polyzoa. We have here again therefore complex and similar organs of diverse and independent origin.
In the highest class of animals (the Mammalia) we have almost always a placental mode of reproduction, i.e. the blood of the fœtus is placed in nutritive relation with the blood of the mother by means of vascular prominences. No trace of such a structure exists in any bird or in any reptile, and yet it crops out again in certain sharks. There indeed it might well be supposed to end, but, marvellous as it seems, it reappears in very lowly creatures; namely, in certain of the ascidians, sometimes called tunicaries or sea-squirts.
Now, if we were to concede that the ascidians were the common ancestors[[61]] of both these sharks and of the higher mammals, we should be little, if any, nearer to an explanation of the phenomenon by means of "Natural Selection," for in the sharks in question the vascular prominences are developed from one fœtal structure (the umbilical vesicle), while in the higher mammals they are developed from quite another part, viz. the allantois.
Upper Figure—Antechinus minutissimus (implacental).