Specimen examined: C. occidentalis californicus, one from Mary's Peak, Benton Co., Oregon, 30, F. W. Sturges' collection.
Clethrionomys glareolus Schreber
Fig. 13
Baculum: Stalk elongate, greatest length (2.9 mm.) twice the greatest breadth in the specimen examined, flattened proximally, greatest length almost 6 times greatest depth of base; three well-developed ossified processes; median process arched in a dorsoventral plane, with basal notch and lateral lobes; lateral processes as long as median process, bowed in dorsal view, flattened distally, with ventromedial vane; basal tuberosities of stalk weakly developed, medially confluent; posterior profile in dorsal view evenly rounded; in end-view dorsal concavity shallow in comparison to most species but deeper than ventral concavity, constriction ¾ greatest depth; shaft straight, at mid-point slightly wider than high, elongate, widest point of stalk less than ¼ of total length from proximal end, slight lateral inflation at tip; lateral profile in dorsal view sloping at first abruptly and then gradually from widest point of stalk anteriorly onto shaft.
The specimen of Clethrionomys glareolus figured by Ognev (1950:31) in dorsal view as I interpret it, resembles my specimen in the rounded base; in the elongate, distally inflated shaft; in the initially abrupt slope of the lateral profile in dorsal view from the greatest width of stalk anteriorly; and in the presence of three well ossified processes. Ognev's specimen differs from mine in the median process being more elongate relative to its width, and rounded proximally, lacking lateral lobes and basal notch; in lateral processes being less curved; in the greater terminal inflation of the shaft; and in the closer approximation of the terminal processes to the shaft. The baculum of Clethrionomys glareolus as described and figured by Didier (1954:243-244) resembles my specimen in general proportions, but is more pointed proximally and more curved in dorsoventral plane. Didier states that the baculum is rather variable in form in this species, in different regions, but that a large number of specimens must be examined to assess the geographic nature of this variation.
Specimen examined: One from Zermatt, Valais, Switzerland, 67100.
Clethrionomys rufocanus Sundevall
Fig. 9
Baculum: Base of stalk broad but relatively flattened dorsoventrally, greatest length (3.2 mm.) less than 1½ greatest width, 4 times greatest depth; three well-developed ossified processes; median process arched in dorsoventral plane, having basal notch and lateral lobes; lateral processes as long as median process, flattened distally, with ventromedial vane; basal tuberosities of stalk weakly developed, medially confluent; posterior profile in dorsal view convex with rounded posterior apex; dorsal surface of base almost flat, ventral concavity broad and shallow; constriction ¾ greatest depth (not including an unusual irregularity on the ventral surface of the base); shaft straight, at mid-point of stalk distinctly wider than high, slender at distal end, widest point of stalk almost 1/3 of total length from proximal end, tip of shaft rounded; lateral profile in dorsal view gradually sloping from widest point anteriorly onto shaft.
The specimen of Clethrionomys rufocanus figured by Ognev (1950:97) resembles my specimen in the presence of three well ossified processes. Ognev's specimen differs however in the lack of a proximal notch on the median process, the lesser proportion of the stalk included in the basal enlargement, the more posterior position of the point of greatest width, and the presence of a concavity in the posterior profile of the stalk in dorsal view. These differences in the stalk may be owing to a difference in age (my specimen perhaps being older).