In the British Islands most of our larger movements are at their start or their finish, or both (so far as our area is concerned), oversea passages, and unless the weather be absolutely favourable, birds do not undertake these voyages. No one has added more to our knowledge of the connection, in what we may term British migration, than Mr Eagle Clarke, but it must not for a moment be imagined that his conclusions and the data from which he arrived at them are purely insular. The British Islands are merely the field of observation, the centre of the field, of the movements of Holarctic birds which travel regularly or occasionally through Britain. Mr Clarke points out repeatedly that in studying the phenomena it is the conditions at the point of departure not at the point of arrival—generally the point of observation—which are important.
The oft-repeated assertion that birds can foretell the nature of approaching weather—that they are living barometers—is not supported by any satisfactory evidence, but it is certain that on many occasions the weather into which they have passed in moving from one zone to another has not only retarded, checked, or exhausted them, but has proved fatally disastrous. During the westward rushes in winter, when exceptionally severe weather has cut off the food-supply of ground-feeding birds, observers who have seen the birds moving in front of the storm have maintained that they had felt its approach and retreated in time. The truth seems to be that the birds start so soon as the supply is cut off but in many cases speedily outstrip the storm. When these exceptional winter migrations take place the birds in the lowlands of Lancashire and Cheshire move westward towards Ireland, and are observed at different points along the North Wales coast. They are sometimes seen travelling in a snow-storm and sometimes in advance of it. In eastern Cheshire I have seen parties of lapwings passing over westward just in advance of snow, which when it reached the East Cheshire fields, started the local lapwings after their relatives from farther east.
During regular migration birds start in favourable weather but frequently meet with unfavourable weather before their arrival at the point aimed at; most of the bird "disasters" at the lighthouses and lightships, and more occasionally inland, can be explained in this way.
In his digest of the observations at lighthouses and lightships Mr Eagle Clarke shows that spells of genial weather are favourable and that during these spells migration is even flowing and continuous [(15)]. Slightly unsettled conditions have little effect, but an increase of the irregularities accelerates migration. Sooner or later cyclonic disturbances interrupt regular movements, and, if these are extraordinary, act as barriers, either holding the birds in one place or forcing a hurried departure or "rush." Favourable weather immediately following a check or "hold up" often causes a rush; a sudden fall in temperature may force large numbers of birds on in autumn or retard them in spring. Temperature, he declares, is the main controlling factor in all extraordinary movements, other meteorological conditions being suitable.
In the autumn migration to Britain, the chief movements take place when a large and well-defined anticyclone has its centre somewhere over Scandinavia, with gentle gradients in a south-westerly direction over the North Sea. Coincident with this we usually find cyclonic conditions prevailing to the west of the British area, with low-pressure centres off the west or south-west of Ireland. The weather is clear and cold, with light variable airs over Scandinavia, but in Britain the sky is overcast, and the wind easterly and moderate to strong; not infrequently these conditions mean fog on our eastern coasts. If the birds leave Scandinavia under favourable conditions they may be met by the approaching cyclonic system, which usually, though by no means always, travels in a north-easterly direction across the Atlantic. Migration is thus checked, but a return of favourable anticyclonic conditions starts the birds again, often with a fresh impulse in the shape of falling temperature. When the anticyclonic area is exceptionally large, extending from the Scandinavian peninsula in a south-westerly direction and embracing the whole of the British Islands, simultaneous immigration and emigration may be witnessed.
Cyclonic spells are not always unfavourable to migration. In spring, when they are of a mild type with soft rain and warm winds following after a cold anticyclonic period, a northward movement is frequent.
Mr Eagle Clarke says that the importance of winds is overstated, but as an incentive only. The direction of the wind has no influence as an incentive but its force is an important factor; in a strong wind a bird may be blown out of its course. Birds will not start in a high wind but may pass into the influence of strong winds which may affect both progress and direction. He adds that particular winds usually prevail during the season of great autumn movements, which are not incentives but are the result of pressure distribution which is favourable to migration. These are usually north-east to south, but a westerly wind would serve as well, but it indicates a pressure distribution which is fatal to migration between north-west Europe and Britain—cyclonic areas to the north-east and east of our area.
All this, no doubt, is perfectly true. It is founded on the analysis of a huge number of carefully recorded observations, and upon a general knowledge of migration which few can ever hope to equal. Mr Clarke understands his subject. It appears, however, to me that he may put rather too much weight upon the barometric influence, and too little on one side of the wind question. Are we yet in a position to say that birds do not make direct use of certain winds? It may be that the use of the prevailing winds at migration time is far more unconsciously intentional (if such an expression can be used) than is at first apparent.
One or two points must be kept well to the front which are often ignored by observers. Firstly, very much visible migration is abnormal; that is to say, most of the incidents of passage which are noticeable, especially observations at the lightships and lighthouses, are during spells of weather which are described as unfavourable; it is the "hold-ups," checks, and "rushes," which attract attention far more than the even-flowing normal migration.
Mr J. Tomison, in his valuable notes on observations made at Skerryvore [(52)], shows that in ordinary clear weather birds pass at a great height, beyond the power of vision. He proves this by instances of the diurnal passage of redwings, birds which are generally supposed to migrate at night, and undoubtedly do so frequently. He heard the well-known passage-note in the daytime, but with the naked eye could see no redwings; he found them with the telescope and later discovered others which were passing above the range of normal vision. Mr Eagle Clarke, commenting upon the extraordinary numbers of rare and exceptional visitors which are noticed on many islands—Fair Island, the Flannens, the Isle of May, and Heligoland may be taken as a few examples—says that it is their detached position and comparatively small size which makes these islands so useful to the observer. The same variety of birds and greater numbers reach larger islands and tracts of land, but they are unobserved when they are thinly distributed and not massed or confined in a small area. "With all our great army of trained observers," he declares, "we in Britain see only an infinitesimal number of the migrants which visit our shores ..." and "this is especially the case on the mainland."