Fig. 10. The left-hand loaf was made from average English wheat. The loaf in the centre was made from Burgoyne’s Fife, and is practically identical in size and shape with the right-hand loaf which was made from imported No. 1 Manitoba
It was at this point that Mendel’s discoveries came to the rescue. Working on the Mendelian lines already explained, Biffen at Cambridge crossed Red Fife with many of the best English varieties. From one of the crosses he was able to isolate a new variety in which are combined the strength of Red Fife and the vigour and cropping power of the English parent. This variety, known as Burgoyne’s Fife, has been grown and distributed by members of the Millers’ Association. In 1911 on the average of 28 separate trials it yielded 38 bushels per acre, which is well above the average of the best English varieties. It has been repeatedly milled and baked, and its strength is between 90 and 100, practically the same as that of Red Fife. It has been awarded many prizes at agricultural shows for quality, and it commands on markets where the local millers have found out its baking qualities about the same price as the best foreign strong wheats, that is to say from 4s. to 5s. per quarter more than the average price of home grown wheat. Taking a fair average yield of wheat as four quarters per acre, Burgoyne’s Fife gives to the farmer an increased return over the ordinary varieties of about 16s. per acre. The introduction of such a variety makes the production of strong wheat in England a practicable reality, and will be a boon both to the farmer and to the inland miller. It is likely too that the possibility of obtaining a better return per acre will induce farmers to grow more wheat. Anything that tends to increase the production of home grown wheat and makes Great Britain less dependent on foreign supplies is a national asset of the greatest value.
It is of the greatest importance to the miller that he should be able to determine the strength of the wheats he buys. Obviously the method mentioned above, which entails milling enough of the sample to enable him to bake a batch of bread, is far too lengthy to be of use in assessing the value of a sample with a view to purchase. The common practice is for the miller or corn merchant to buy on the reputation of the various grades of wheat, which he confirms by inspection of the sample. Strength is usually associated with certain external characters which can readily be judged by the eye of the practised wheat buyer. Strong wheats are usually red in colour, their skin is thin and brittle, the grain is usually rather small, and has a very characteristic horny almost translucent appearance. The grains are extremely hard and brittle, and when broken the inside looks flinty. On chewing a few grains the starch is removed and there remains in the mouth a small pellet of gluten, which is tough and elastic like rubber, but not sticky.
Weak wheats as a rule possess none of these characters. Their colour may be either red or white, their skin is commonly thick and tough, the grain is usually large and plump, and often has an opaque mealy appearance. It is soft and breaks easily, and the inside is white, soft and mealy. Very little gluten can be separated from it by chewing, and that little is much less tough and elastic than the gluten of a strong wheat.
These characters, however, are on the whole less reliable than the reputation of the grade of wheat under consideration. To make a reliable estimate of strength from inspection of a sample of wheat requires a natural gift cultivated by continual practice. Even the best commercial judges of wheat have been known to be deceived by a sample of white wheat which subsequent milling and baking tests showed to possess the highest strength. The mistake was no doubt due to the great rarity of strength among white wheats. This rarity will doubtless soon disappear now that a pure strain of White Fife has been isolated and shown to possess strength quite equal to that of Red Fife. Sometimes too the ordinary home grown varieties produce most deceptive samples which show all the external characters of strong wheats. Such samples, however, on milling and baking are invariably found to possess the usual strength of home grown wheat, about 65 on the scale. These considerations show the great need of a scientific method of measuring strength, which can be carried out rapidly and on a small sample of grain. This need is felt at the present time not only by the miller and the merchant, but by the wheat breeder. For instance, in picking out the plants possessing strong grain from cultures of the second generation after making his crosses, the plant breeder up to the present has had to rely on inspection by eye, and on the separation of gluten by chewing, for a single plant obviously cannot yield enough grain to mill and bake. This fact no doubt explains the differences of opinion among plant breeders on the inheritance of strength, for it is not every one who can acquire the power of judging wheat accurately by his senses. Such a faculty is a personal gift, and is at best apt to fail at times.
The search for a rapid and accurate method of measuring strength has for many years attracted the attention of investigators. As might be expected most of the investigations have centred round the gluten, for as mentioned above the gluten of a strong wheat is much more tough and elastic than that of a weak wheat. Gluten is a characteristic constituent of all wheats, and it is the presence of gluten which gives to wheat flour the power of making bread. The other cereals, barley, oats, maize and rice are very similar to wheat in their general chemical composition, but they do not contain gluten. Consequently they cannot make bread.
In making bread flour is mixed with water and yeast. The yeast feeds on the small quantity of sugar contained in the flour, fermenting it and forming from it alcohol and carbon dioxide gas. The gluten being coherent and tough is blown into numberless small bubbles by the gas, which is thus retained inside the bread. On baking, the high temperature of the oven fixes these bubbles by drying and hardening their walls, and the bread is thus endowed with its characteristic porous structure. If a cereal meal devoid of gluten is mixed with water and yeast, fermentation will take place with formation of gas, but the gas will escape at once, and the product will be solid and not porous. Evidently from the baking point of view gluten is of the greatest importance. One of the most obvious methods that have been suggested for estimating the strength of wheat depends on the estimation of the percentage of gluten contained in the flour. The method has not turned out very successfully, for strength seems to depend rather on the quality than on the quantity of gluten in the wheat. Much attention has been given to the study of the causes of the varying quality of the gluten of different wheats. Gluten for instance has been shown to be a mixture of two substances, gliadin and glutenin, and the suggestion has been made that its varying properties are dependent on the varying proportions of these two substances present in different samples. This suggestion however failed to solve the problem.
After seven years of investigation the author has worked out the following theory of the strength of wheat flours, which has finally enabled him to devise a method which promises to be both accurate and rapid, and to require so little flour that it can readily be used by the wheat breeder to determine the strength of the grain in a single ear. It has already been mentioned that a strong wheat is one that will make a large loaf of good shape and texture. The strength of a wheat may therefore be defined as the power of making a large loaf of good shape and texture. Evidently strength is a complex of at least two factors, size and shape, which are likely to be quite independent of each other. Not infrequently, for instance, wheats are met with which make large loaves of bad shape, or on the other hand, small loaves of good shape. Probably therefore the size of the loaf depends on one factor, the shape on another; and the failure of the many attempts to devise a method of estimating strength have been caused by the impossibility of measuring the product of two independent factors by one measurement.
It seemed a feasible idea that the size of the loaf might depend on the volume of gas formed when yeast was mixed with different flours. On mixing different flours with water and yeast it was found that for the first two or three hours they all gave off gas at about the same rate. The reason of this is that all flours contain about the same amount of sugar, approximately one per cent., so that at the beginning of the bread fermentation all flours provide the yeast with about the same amount of sugar for food. But this small amount of sugar is soon exhausted, and for its subsequent growth the yeast is dependent on the transformation of some of the starch of the flour into sugar. Wheat like many other seeds contains a ferment or enzyme called diastase, which has the power of changing starch into sugar, and the activity of this ferment varies greatly in different wheats. The more active the ferment in a flour the more rapid the formation of sugar. Consequently the more rapidly the yeast will grow, and the greater will be the volume of gas produced in the later stages of fermentation in the dough. As a rule it is not practicable to get the dough moulded into loaves and put into the oven before it has been fermenting for about six or eight hours. If the flour possesses an active ferment it will still be rapidly forming gas at the end of this time, and the loaf will go into the oven distended with gas under pressure from the elasticity of the gluten which forms the walls of the bubbles. The heat of the oven will cause each gas bubble to expand, and a large loaf will be the result. If the ferment of the flour is of low activity it will not be able to keep the yeast supplied with all the sugar it needs, the volume of gas formed in the later stages of the fermentation of the dough will be small, the dough will go into the oven without any pressure of gas inside it, little expansion will take place as the temperature rises, and a small loaf will be produced.
From these facts it is quite easy to devise a method of estimating how large a loaf any given flour will produce. The following method is that used by the author. A small quantity of the flour, usually 20 grams, is weighed out and put into a wide mouthed bottle. A flask of water is warmed to 40° C., of this 100 c.c. is measured out, and into it 2½ grams of compressed yeast is intimately mixed, 20 c.c. of the mixture being added to the 20 grams of flour in the bottle. The flour and yeast-water are then mixed into a cream by stirring with a glass rod. The bottle is then placed in a vessel of water which is kept by a small flame at 35° C. The bottle is connected to an apparatus for measuring gas, and the volume of gas given off every hour is recorded. As already mentioned all flours give off about the same volume of gas during the first three hours. After this length of time the volume of gas given off per hour varies greatly with different flours. Thus a flour which will bake a large loaf gives off under the conditions above described about 20 c.c. of gas during the sixth hour of fermentation, whilst a flour which bakes a small tight loaf gives off during the sixth hour of fermentation only about 5 c.c. of gas.