Two other grass-eating beasts, of large size—although smaller than the above—were also plentiful. One, a bison, bigger, straighter-horned and less specialized than our modern bison, represented the cattle, which were among the animals that passed to America over the Alaskan land bridge in Pleistocene time.
The other was a big, coarse-headed horse, much larger than any modern wild horse, and kin to the then existing giant horse of Texas, which was the size of a percheron. The horses, like the camels, had gone through their developmental history on this continent, the earliest ancestor, the little four-toed "dawn horse" of the Eocene, being likewise the size of a jack-rabbit. Through millions of years, while myriads of generations followed one another, the two families developed side by side, increasing in size and seemingly in adaptation to the environment. Each stock branched into many different species and genera. They spread into the Old World and into South America. Then, suddenly,—that is, suddenly in zoologic sense—both completely died out in their ancient home, and the horses in South America also, whereas half a dozen very distinct species are still found in Asia and Africa.
All these great creatures wandered in herds to and fro across the grassy Californian plains and among the reaches of open forest. Preying upon them were certain carnivores grimmer and more terrible than any now existing. The most distinctive and seemingly the most plentiful was the sabretooth. This was a huge, squat, short-tailed, heavily built cat with upper canines which had developed to an almost walrus-like length; only, instead of being round and blunt like walrus tusks, they were sharp, with a thin, cutting edge, so that they really were entitled to be called sabres or daggers. Whether the creature was colored like a lion or like a tiger or like neither, we do not know, for it had no connection with either save its remote kinship with all the cats. The sabretooth cats, like the true cats, had gone through an immensely long period of developmental history in North America, although they did not appear here as early as the little camels and horses. Far back across the ages, at or just after the close of the Eocene—the "dawn age" of mammalian life—certain moderate-sized or small cat-like creatures existed on this continent, doubtless ancestral to the sabretooth, but so generalized in type that they display close affinities with the true cats, and even on certain points with the primitive dog creatures of the time. Age followed age—Oligocene, Miocene, Pliocene. The continents rose and sank and were connected and disconnected. Vast lakes appeared and disappeared. Mountain chains wore down and other mountain chains were thrust upward. Periods of heat, during which rich forests flourished north of the arctic circle, were followed by periods of cold, when the glacial ice-cap crept down half-way across the present temperate zone. Slowly, slowly, while the surface of the world thus changed, and through innumerable reaches of time, the sabretooth cats and true cats developed along many different lines in both the Old World and the New. One form of sabretooth was in Europe with the bestial near-human things who were the immediate predecessors of the first low but entirely human savages. It was in the two Americas, however, that the sabretooth line culminated, immediately before its final extinction, in its largest and most formidable forms. This California sabretooth was not taller than a big cougar or leopard, but was probably as heavy as a fair-sized lion. Its skeletal build is such that it cannot have been an agile creature, apt at the pursuit of light and swift prey. By rugged strength and by the development of its terrible stabbing and cutting dagger teeth, that is, by sheer fighting ability, it was fitted for attack upon and battle with the massive herbivores then so plentiful. It must indeed have been a fearsome beast in close grapple. Doubtless with its sharp, retractile claws it hung onto the huge bodies of elephant, camel, and ground-sloth, of horse and bison, while the sabres were driven again and again into the mortal parts of the prey and slashed the flesh as they withdrew. It seems possible that the mouth was opened wide and stabbing blows delivered, almost as a rattlesnake strikes with raised fangs. Vast numbers of sabretooth skeletons have been found in the asphalt; evidently the strange, formidable creature haunted any region which held attraction for the various kinds of heavy game on which it preyed.
The only other carnivore as abundant as the sabretooth was a giant wolf. This was heavier than any existing wolf, with head and teeth still larger in proportion. The legs were comparatively light. Evidently, like the sabretooth, this giant wolf had become specialized as a beast of battle, fitted to attack and master the bulky browsers and grazers, but not to overtake those that were smaller and swifter. The massive jaws and teeth could smash heavy bones and tear the toughest hide; and a hungry pack of these monsters, able to assail in open fight any quarry no matter how fierce or powerful, must have spread dire havoc and dismay among all things that could not escape by flight.
There were two still larger predatory species, which were much less plentiful than either the wolf or the sabretooth. One was a short-faced cave-bear, far larger than even the huge Alaskan bear of to-day. Doubtless it took toll of the herds; but bears are omnivorous beasts, and not purely predatory in the sense that is true of those finished killers, the wolves and big cats. Unlike the wolves and cats, bears were geologically recent immigrants to America.
The other was a true cat, a mighty beast; bigger than the African lion of to-day; indeed, perhaps the biggest and most powerful lion-like or tiger-like cat that ever existed. Seemingly it was much rarer than the sabretooth; but it is possible that this seeming rarity was due to its not lurking in the neighborhood of pools and licks but travelling more freely over the wastes, being of a build fit not only for combat but for an active and wandering life. It is usually spoken of as kin to the African lion, a decidedly smaller beast. It is possible that its real kinship lies with the tiger. The Manchurian form of the tiger is an enormous beast, and a careful comparison of the skulls and skeletons may show that it equals in size the huge western American cat of Pleistocene times. I have already spoken of the fact that in many cases it is almost impossible to distinguish the lion and tiger apart by the bones alone; and it may be that the exact affinities of these recently extinct species with living forms cannot be definitely determined. But during historic and prehistoric times the lion has been a beast of western Eurasia and of Africa. The tiger, on the contrary, is and has been a beast of eastern Asia, and apparently has been spreading westward and perhaps southward—that it was not as ancient an inhabitant of jungle-covered southern India as the elephant and leopard seems probable from the fact that it is not found in Ceylon, which island in all likelihood preserves most of the southern Indian fauna that existed prior to its separation from the mainland. Moreover, the finest form of tiger exists in cold northeastern Asia. In Pleistocene times this portion of Asia was connected by a broad land bridge with western America, where the mighty American cat then roved and preyed on the herds of huge plant-eating beasts. We know that many Asiatic beasts crossed over this land bridge—the bears, bison, mountain-sheep, moose, caribou, and wapiti, which still live both in Asia and North America, and the mammoth and cave-bears, which have died out on both continents. It is at least possible—further investigation may or may not show it to be more than possible—that the huge Pleistocene cat of western America was the collateral ancestor of the Manchurian tiger. Whether it was another immigrant from Asia, or a developed form of some big American Pliocene cat, cannot with our present knowledge be determined.
Surely the thought of this vast and teeming, and utterly vanished wild life, must strongly appeal to every man of knowledge and love of nature, who is gifted with the imaginative power to visualize the past and to feel the keen delight known only to those who care intensely both for thought and for action, both for the rich experience acquired by toil and adventure, and for the rich experience obtained through books recording the studies of others.
Doubtless such capacity of imaginative appreciation is of no practical help to the hunter of big game to-day, any more than the power to visualize the long-vanished past in history helps a practical [politician] to do his ordinary work in the present workaday world. The governor of Gibraltar or of Aden, who cares merely to do his own intensely practical work, need know nothing whatever about any history more ancient than that of the last generation. But this is not true of the traveller. It is not even true of the politician who wishes to get full enjoyment out of life without shirking its duties. He certainly must not become a mere dreamer, or believe that his dreams will help him in practical action. But joy, just for joy's sake, has its place too, and need in no way interfere with work; and, of course, this is as true of the joy of the mind as of the joy of the body. As a man steams into the Mediterranean between the African coast and the "purple, painted headlands" of Spain, it is well for him if he can bring before his vision the galleys of the Greek and Carthaginian mercantile adventurers, and of the conquering Romans; the boats of the wolf-hearted Arabs; the long "snakes" of the Norse pirates, Odin's darlings; the stately and gorgeous war craft of Don John, the square-sailed ships of the fighting Dutch admirals, and the lofty three-deckers of Nelson, the greatest of all the masters of the sea. Aden is like a furnace between the hot sea and the hot sand; but at the sight of the old rock cisterns, carved by forgotten hands, one realizes why on that coast of barren desolation every maritime people in turn, from the mists that shroud an immemorial antiquity to our own day of fevered materialistic civilization, has seized Aden Bay—Egyptian, Sabean, Byzantine, Turk, Persian, Portuguese, Englishman; and always, a few miles distant, in the thirsty sands, the changeless desert folk have waited until pride spent itself and failed, and the new power passed, as each old power had passed, and then the merciless men of the waste once more claimed their own.
Gibraltar and Aden cannot mean to the unimaginative what they mean to the men of vision, to the men stirred by the hero tales of the past, by the dim records of half-forgotten peoples. These men may or may not do their work as well as others, but their gifts count in the joy of living. Enjoyment the same in kind comes to the man who can clothe with flesh the dry bones of bygone ages, and can see before his eyes the great beasts, hunters and hunted, the beasts so long dead, which thronged the Californian land at a time when in all its physical features it had already become essentially what it still continues to be.
The beast life of this prehistoric California must be called ancient by a standard which would adjudge the Egyptian pyramids and the Mesopotamian palace mounds and the Maya forest temples to be modern. Yet when expressed in geologic terms it was but of yesterday. When it flourished the Eurasian hunting savages were in substantially the same stage of progress as the African hunting savages who now live surrounded by a similar fauna. On the whole, taking into account the number, variety, and size of the great beasts, the fauna which surrounded Palæolithic man in Europe was inferior to that amid which dwell the black-skinned savages of equatorial Africa. Even Africa, however, although unmatched in its wealth of antelopes, cannot quite parallel, with its lion, elephant, and zebras, the lordlier elephant, the great horse, and the huge cat of the earlier Californian fauna; and the giraffe, the hyena, the rhinoceros, and the hippopotamus do not quite offset the sabretooth, the giant wolf, the mastodon, the various species of enormous ground-sloths, and the huge camel; the bison and buffalo about balance each other.