Wooded Stems are divided into two great and well-defined classes, according to their internal conformation, viz. such as grow from without (exogenous), and such as enlarge from within (endogenous). The former are more common in cold, and the latter in hot climates.
Exogenous Stems.—On examining a section of a stem of an oak, or any other of our forest trees, we observe the following parts: first, the pith, or its remains in the centre; second, the bark on the outside; third, a mass of wood between the two, broken up into portions by the concentric deposition of the layers, and by a series of lines which pass from the centre to the circumference. Thus, there are always pith, bark, wood, and medullary rays. Each stem has two systems, the cellular or horizontal, and the vascular or longitudinal, and the parts just mentioned must belong to one or other of those systems. Thus, the pith, medullary rays, and bark belong to the horizontal system; and the wood constitutes the longitudinal system.
The Pith occupies the centre of the stem, and remains throughout the period of growth of some trees, as of the elder; or is abstracted after a few years, as in the oak, and almost all large trees. In the latter class of trees, there are some remains of the pith for many years after the process of absorption has commenced, but at length no vestige can be detected, and its position is known only by the central spot around which the wood is placed in circles. In the old age of the tree the pith frequently assumes a colour which it has obtained from the juices which have been deposited. The connections of the pith are extremely important. Firstly, it is in direct connection with every branch, and is the structure which first conveys fluids to, and receives fluids from every new leaf. It thence becomes the main organ of nutriment, and, at the same time, the chief depository of the secretions. Secondly, it is in equally direct and unbroken connection with the bark, through the medium of the medullary rays; and so becomes the centre of all the movements of sap which proceed in the horizontal system.
The mode in which the ultimate disappearance of the pith occurs has been a matter of speculation. That the circulation in the heart-wood ceases after a certain number of years, and that the connection between it and the bark becomes broken, is proved by the fact that numbers of trees may be found in tolerably vigorous growth within the bark, whereas at the heart they are decayed and rotten. It appears clear that it is not converted into wood, and there are facts against the opinion that it is gradually compressed by the wood; but since it is known that in the growth of the tree much compression of the previously formed wood must occur, and since this compression is a likely theory by which to account for the disappearance of the less resisting pith, it is now generally considered to be one of the causes of this occurrence. As a general rule, the pith, so long as it exists, is not mingled with other than cellular structures; but, in certain instances, wooden fibre has been found with it, and, in others, spiral vessels have been detected.
Medullary Sheath.—Immediately surrounding the pith of all exogenous plants, there is a layer of longitudinal tissue, which has received the name of medullary sheath. This sheath has no special walls, but is bounded by the pith on the inner, and the wood on the outer side. It is in this situation that ducts of various kinds and spiral vessels may be found, and in all cases it conveys the longitudinal structure from the root, direct to each leaf. The integrity of this structure is therefore highly necessary to the life of the tree.
Medullary Rays.—These structures come next in order, and, as has been previously intimated, belong to the horizontal cellular system of the stem; they constitute the channel of communication between the bark and the pith, and are composed of a series of walls of single cells resting upon the root, and proceeding to the top of the tree, and radiating from the centre. They lie between the wedge-like blocks of wood, and as they have a lighter colour than the wood, they are evident on an oblique section of any stem, and are called the silver grain. Their colour and number suffice to enable anyone to distinguish various kinds of wood, and greatly increase their beauty. They cannot, of course, exist before the wood is formed, and are therefore not met with in very young trees. They commence to exist with the first deposited layers of wood, and continue to grow outwardly, or nearest to the bark, so long as the wood continues to be deposited. In those woods which possess in abundance the silver grain, another source of ornament exists, viz. a peculiar damask or dappled effect, somewhat similar to that artificially produced on damask linens, moreens, silks, and other fabrics, the patterns on which result from certain masses of the threads on the face of the cloth running lengthways, and other groups crossways. This effect is observable in a remarkable degree in the more central planks of oak, especially in Dutch wainscot.
The Bark.—As the medullary rays terminate in the bark, on their outer side, the consideration of that part next follows. It forms the sheath of the tree, and its more immediate use is that of giving protection to the wood. If bark did not exist, there would be no formative fluid, and without formative fluid there could not be any deposit of woody fibre.
Wood.—We find wood occupying nearly the whole body of the trunk of the tree, and arranged, as a rule, in a very regular manner. On taking up any piece of wood, but more particularly the entire section of a stem, we first notice a series of circles, which increase in diameter and separate by wider intervals as we approach the bark. In this manner the trunk is composed of numerous zones enclosed within each other. Again, in almost all trees, the medullary rays before mentioned may be observed passing in straight lines from the centre to the circumference; and, as the circle of the stem at the bark is much larger than any circle near to the centre, it follows that the medullary rays will be wider apart at the bark than at the pith. On this view of the subject it may be stated that the stem is composed of a series of wedge-shaped blocks, which have their edges meeting at the centre. The combination of these two views gives the correct idea of the arrangement of the wood, viz. a series of wedges, each divided into segments of unequal width by circular lines passing across them. From this description it must not be imagined that these various portions are detached from each other; for although the medullary rays and the circular mode of deposition both tend to a less difficult cleavage of the wood, they yet bind the parts very closely to each other.
The explanation of the occurrence of distinct zones of wood is, that each zone is the produce of one year, and that in our climate, more so than in tropical climates, the period of growth of wood ceases for many months between the seasons, and this induces a distinction in appearance between the last wood of a former, and the first wood of a succeeding year. This distinction is maintained throughout each year, and throughout a long series of years.
The enclosure of zone within zone, is owing to the mode in which the wood is produced, and the position in which it is deposited. Wood is formed by the leaves during the growing season, and passes down towards the root between the bark and the wood of the previous year; and, as the leaves more or less surround the whole stem, the new layer at length completes a zone, and perfectly encloses the wood of all former years. This is the explanation of the term exogenous, which is derived from two words signifying to grow, outwardly, for the stem increases in thickness by successive layers on the outer side of the previously formed wood.