This external part of the ear is differently formed in different animals; and admirably suited to their various situations and habits. In man it is close to the head, but so formed as to collect the various pulses with great accuracy; in other animals it is more simple, where less accuracy is required, but it is, in general, much larger, having the appearance of an oblong funnel; and this gives them a greater delicacy of hearing, which was necessary for them.
In animals which are defenceless and timid, and which are constantly obliged to seek their safety in flight, the opening of this funnel is placed behind, that they may better hear the noises behind them. This is particularly instanced in the hare. Beasts of prey have this opening before, that they may more easily discover their prey; as the lion and tiger. Those that feed on birds have the opening directed upwards, as the fox; and it is inclined downwards in animals, such as the weasel, which seek their prey on the earth.
To this external part of the ear, which I have described, is connected the meatus auditorius, or passage to the internal ear, which is somewhat of a compressed cylindrical figure, lessening as it bends inwards: a considerable part of it is bony, and it is bent towards the middle. Across this passage, at its inner extremity, is stretched a thin membrane, called membrana tympani. Upon the surface of this membrane, the sonorous waves, which have been directed inwards by the external ear, strike, and cause it to vibrate like the membrane of a drum. This membrane is stretched over a cavity in the bone, called the os petrosum, which cavity is called the tympanum, or drum of the ear, which is of a rounded figure, divided in its middle by a promontory, and continued backwards to the cells of the mastoid bone. Besides this continuation of the tympanum into the mastoid cells, it has a free communication with the mouth, by means of a tube I shall soon describe.
Within this cavity of the tympanum are placed four small bones, which facilitate the hearing: the first is the malleus or hammer, so called from its shape: the upper part of its round head rests upon the concavity of the tympanum, from whence the handle is extended down, along the membrane of the tympanum; this bone has several muscles, which move it in different directions, and cause it to stretch or brace the membrana tympani, when we wish to hear with accuracy.
Connected with the malleus is another small bone, called the incus, or anvil, which is connected with another, called the stapes, or stirrup, from its shape. These two bones are connected by a small oval shaped bone, called os orbiculare, placed between them: the whole forming a little chain of bones.
The stapes, or stirrup, has its end of an oval shape, which fits a small hole called fenestra ovalis, in that part of the ear called the labyrinth, or innermost chamber of the ear.
The labyrinth consists of three parts; first, the vestibule, which is a round cavity in a hard part of the os petrosum; secondly, the semicircular canals, so called from their shape, which however is not exactly semicircular; thirdly, the cochlea, which is a beautifully convoluted canal, like the shell of a snail. This part has a round cavity called fenestra rotunda, which is covered with a thin elastic membrane, and looks into the tympanum.
The vestibule, semicircular canals, and cochlea, the whole of which is called the labyrinth, form one cavity, which is filled with a very limpid fluid resembling water, and the whole lined with a fine delicate membrane, upon which the auditory nerve is expanded, like the retina upon the vitreous humor of the eye. This beautiful apparatus was only lately discovered by an Italian physician, Scarpa. The auditory nerve is a portion of the seventh pair, which is called the portio mollis or soft portion.
There is one part of the ear still to be described, namely, the Eustachian tube, so called from Eustachius, the anatomist, who first described it. This tube opens by a wide elliptical aperture into the tympanum behind the membrane; the other end, which gradually grows wider, opens into the cavity of the mouth. By this canal the inspired air enters the tympanum to be changed and renewed, it likewise serves some important purpose in hearing, with the nature of which we are yet unacquainted. It is certain that we can hear through this passage, for if a watch be put into the mouth, and the ears stopped, its ticking may be distinctly heard; and in several instances of deafness, this tube has been found completely blocked up.
The waves, which have been described as propagated in the air, in all directions from the sounding body, enter the external cartilaginous part of the ear, which, as has before been observed, is admirably fitted for collecting and condensing them. As soon as these pulses excite tremors in the membrane of the tympanum, its muscles stretch and brace it, whence it becomes more powerfully affected by these impulses. It is on this account that we hear sounds more distinctly when we attend to them, the membrane being then stretched.