The shape of the eye is nearly spherical; it is composed of several coats or tunics, one within another; and is filled with transparent humours of different densities.
The proper coats of the eye are reckoned five in number; viz. the sclerotica, cornea, choroides, iris or uvea, and the retina.
After the tunica conjunctiva, or adnata, (a membrane, which, having lined the eyelids in the manner of a cuticle, surrounds the anterior part of the globe) is removed, we perceive a white, firm, membrane, called the sclerotica, which takes its rise from that part of the globe where the optic nerve enters, and surrounds the whole eye, except a little in the fore part; which fore part has a membrane, immediately to be described, called the cornea. The tunica sclerotica, viewed through the conjunctiva, forms what is called the white of the eye. Some anatomists have supposed that this coat is a continuation of the dura mater, which surrounds the optic nerve; but later observations have shown this opinion to be ill founded. The tunica sclerotica consists of two layers, which are with difficulty separated.
The next coat is the cornea, so called from its resemblance to transparent horn; it arises where the sclerotic coat ends, and forms the fore part of the eye. The cornea is a segment of a lesser sphere than the rest of the eye, and consequently makes it more prominent on the fore part: it is transparent, and firmly connected by its edges to the sclerotica.
Immediately adherent to the sclerotica, within, is the choroides, which takes its rise from that part of the eye where the optic nerve enters, and accompanies the sclerotica to the place where it is joined to the cornea; here it is very closely connected to the sclerotica, where it forms that annulus, called ligamentum ciliare; then leaving the sclerotic coat, it is turned inwards, and surrounds the crystalline lens; but as this circle, where it embraces the crystalline, is much narrower than where the membrane leaves the sclerotic coat, it becomes beautifully corrugated, which folds or corrugations have been, by the more ancient anatomists, improperly called ciliary processes.
To the same part of the choroid coat, where the ciliary ligament begins, is fixed a moveable and curious membrane, called the iris; this membrane has a perforation in the middle, called the pupil, for the admission of the rays of light. The iris is composed of two kinds of fibres: those of the one sort tend, like the radii of a circle, towards its centre, and the others form a number of concentric circles round the same centre. The pupil is of no constant magnitude, for when a very luminous object is viewed, the circular fibres of the iris contract, and diminish its orifice; and, on the contrary, when objects are dark and obscure, those fibres relax, and suffer the pupil to enlarge, in order to admit a greater quantity of light into the eye: it is thought that the radial fibres also assist in enlarging the pupil. The iris is variously coloured in different persons, but according to no certain rule; though in general, they who have light hair, and a fair complexion, have the iris blue or grey; and, on the contrary, they whose hair and complexion are dark, have the iris of a deep brown; but whether this difference in colour occasions any difference in the sense, is not yet discovered. In the human eye the whole choroid coat, and even the interior surface of the iris or uvea, is lined with a black mucus; this mucus, or as it is called, pigmentum, is darkest in young persons, and becomes more light coloured as we advance in years. In many animals, but more particularly those which catch their prey in the night, this pigmentum is of a bright colour: its use will appear afterwards.
The last, and innermost coat of the eye, is the retina, it differs much from the above mentioned coats, being very delicate and tender. It is nothing but an expansion of the medullary part of the optic nerve, which is inserted into each eye, nearer the nose, and a little higher, than the axis. This coat has been thought by many to end where the choroides, going inwards, towards the axis of the eye, forms the ciliary ligament; Dr. Monro thinks that it is not continued so far, and we cannot see with what advantage it could have been continued to the ciliary ligament, since none of the rays of light, passing through the pupil, could fall upon that part of it. In the middle of the optic nerve is found the branch of an artery, from the internal carotid, which is diffused and ramified in a beautiful manner along the retina. From this artery, a small branch goes through the middle of the vitreous humour, and giving off branches on every side, expands itself upon the capsule of the crystalline lens.
We shall now consider the humours of the eye, which are three in number, the aqueous, the crystalline, and the vitreous; all transparent, and in general colourless; but of different densities.
The aqueous humour, so called from its resemblance to water, fills up all the space between the cornea and the crystalline humour. It is partly before and partly behind the uvea, and is divided by that membrane into two parts, which are called the chambers of the aqueous humour; which chambers communicate with each other by means of the pupil.
The next humour is the crystalline; it is situated between the aqueous and vitreous humours, and is connected to the choroid coat by the ciliary ligament: it is not the least of all the humours, as has been generally supposed, the aqueous and it being of equal weights; but its substance is more firm and solid than that of the other humours: its figure is that of a double convex lens; but the fore part next the pupil is not so convex as its other side, which is contiguous to the vitreous humour; the diameter of the sphere, of which its anterior segment is a part, being in general about seven or eight lines, whereas the diameter of the sphere, of which its posterior segment forms a portion, is commonly only about five or six lines. It is covered with a fine transparent capsule, which is called arachnoides. This humour is situated exactly behind the pupil, but not in the centre of the eye, as was supposed by Vesalius, being a good deal nearer its forepart. The convexity of its posterior surface is received into an equal concavity of the vitreous humour. It is not of an equal density throughout, but is much more hard and dense towards its centre than externally, the reason of which will appear hereafter. Till we arrive at about our thirtieth year, this humour continues perfectly transparent, and colourless; about that time it generally has a little tinge of yellow, and this colour increases with age.